U. zammaranoi is hypothesized to be the sister group to Heteropneustes and the remaining clariids. The problematic relationship of this species to the rest of the family is due to the lack of a labyrinth organ that characterizes the family.[4]
^Devaere, Stijn; Adriaens, Dominique; Teugels, Guy G.; Verraes, Walter (2005). "Morphology and spatial constraints in a dorsoventrally flattened skull, with a revised species description of Platyallabes tihoni (Poll, 1944)". Journal of Natural History. 39 (19): 1653–1673. CiteSeerX10.1.1.596.7003. doi:10.1080/00222930400020103. S2CID59363346.
^Romero, A., editor (2001). The Biology of Hypogean Fishes. Developments in Environmental Biology of Fishes. ISBN978-1402000768