List of crocidurines
Species in mammal subfamily Crocidurinae
Greater white-toothed shrew (Crocidura russula )
Crocidurinae is a subfamily of small mammals in the shrew family Soricidae , which in turn is part of the order Eulipotyphla . A member of this family is called a crocidurine, or a white-toothed shrew. Crocidurinae is one of three subfamilies in Soricidae, along with the African shrews of Myosoricinae and the red-toothed shrews of Soricinae . They are found in Africa, Europe, and Asia, primarily in forests, savannas , shrublands , and grasslands , though some species can also be found in wetlands, rocky areas, or coastal areas. They range in size from the Etruscan shrew , one of the smallest mammal species at 3 cm (1 in) plus a 2 cm (1 in) tail,[ 1] to the hero shrew and Thor's hero shrew , at 15 cm (6 in) plus a 10 cm (4 in) tail. Crocidurines primarily eat invertebrates , small mammals, frogs, toads, and lizards. Almost no crocidurines have population estimates, but 21 species are categorized as endangered species , while the Andaman shrew , Christmas Island shrew , Harenna shrew , Jenkins's shrew , Nicobar shrew , and Wimmer's shrew are categorized as critically endangered .
The 235 extant species of Crocidurinae are divided into nine genera ; 192 of them are in Crocidura , the most species of any mammal genus.[ 2] A few extinct prehistoric Crocidurinae species have been discovered, though due to ongoing research and discoveries the exact number and categorization is not fixed.[ 3]
Conventions
Conservation status codes listed follow the International Union for Conservation of Nature (IUCN) Red List of Threatened Species . Range maps are provided wherever possible; if a range map is not available, a description of the crocidurine's range is provided. Ranges are based on the IUCN Red List for that species unless otherwise noted.
Classification
The subfamily Crocidurinae consists of nine genera: Crocidura , containing 192 species; Diplomesodon and Feroculus with a single species each; Paracrocidura , with three species; Ruwenzorisorex , with one species; Scutisorex , containing two species; Solisorex , with one species, Suncus , containing nineteen species; and Sylvisorex , containing fifteen species.
Subfamily Crocidurinae
Genus Crocidura (white-toothed shrews): 191 species
Genus Diplomesodon (piebald shrew): one species
Genus Feroculus (Kelaart's long-clawed shrew): one species
Genus Paracrocidura (large-headed shrews): three species
Genus Ruwenzorisorex (Ruwenzori shrew): one species
Genus Scutisorex (hero shrews): two species
Genus Solisorex (Pearson's long-clawed shrew): one species
Genus Suncus (musk shrews and pygmy shrews): nineteen species
Genus Sylvisorex (forest shrews): fifteen species
Crocidurines
The following classification is based on the taxonomy described by the reference work Mammal Species of the World (2005), with augmentation by generally accepted proposals made since using molecular phylogenetic analysis , as supported by both the IUCN and the American Society of Mammalogists .[ 5]
Genus Crocidura – Wagler , 1832 – 192 species
Common name
Scientific name and subspecies
Range
Size and ecology
IUCN status and estimated population
African black shrew
C. nigrofusca Matschie , 1895
Central and eastern Africa
Size : 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[ 6] Habitat : Forest[ 7] Diet : Invertebrates , small mammals, frogs, toads, and lizards[ 6]
LC
Unknown [ 7]
African dusky shrew
C. caliginea Hollister , 1916
Central Africa
Size : 6–8 cm (2–3 in) long, plus 4–6 cm (2 in) tail[ 8] Habitat : Forest[ 9] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
LC
Unknown [ 9]
African giant shrew
C. olivieri (Lesson , 1827)
C. o. anchietae
C. o. bueae
C. o. cara
C. o. cinereoaenea
C. o. darfurea
C. o. giffardi
C. o. guineensis
C. o. hansruppi
C. o. hedenborgiana
C. o. kivu
C. o. manni
C. o. martiensseni
C. o. nyansae
C. o. occidentalis
C. o. odorata
C. o. olivieri
C. o. spurelli
C. o. sururae
C. o. toritensis
C. o. zuleika
Sub-Saharan Africa
Size : 11–14 cm (4–6 in) long, plus 8–10 cm (3–4 in) tail[ 10] Habitat : Forest, savanna , and shrubland [ 11] Diet : Invertebrates[ 10]
LC
Unknown [ 11]
Andaman shrew
C. andamanensis Miller , 1902
South Andaman Island in India
Size : 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[ 6] Habitat : Forest[ 12] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
CR
Unknown [ 12]
Andaman spiny shrew
C. hispida Thomas , 1913
South Andaman Island in India
Size : 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[ 6] Habitat : Forest[ 13] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
VU
Unknown [ 13]
Annamite shrew
C. annamitensis Jenkins , Lunde , & Moncrieff , 2009
Vietnam
Size : 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[ 6] Habitat : Forest[ 14] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
DD
Unknown [ 14]
Ansell's shrew
C. ansellorum Hutterer & Dippenaar , 1987
South-central Africa
Size : 5–6 cm (2–2 in) long, plus 4–6 cm (2 in) tail[ 15] Habitat : Forest[ 16] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
EN
Unknown [ 16]
Arabian shrew
C. arabica Hutterer , 1988
Oman and Yemen
Size : 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[ 6] Habitat : Unknown[ 17] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
LC
Unknown [ 17]
Armenian shrew
C. armenica Gureev , 1963
Armenia
Size : 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[ 6] Habitat : Rocky areas[ 18] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
DD
Unknown [ 18]
Asian gray shrew
C. attenuata H. Milne-Edwards , 1872
Eastern and southeastern Asia
Size : 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[ 6] Habitat : Forest, shrubland, and grassland [ 19] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
LC
Unknown [ 19]
Asian lesser white-toothed shrew
C. shantungensis Miller , 1901
C. s. quelpartis
C. s. shantungensis
Eastern Asia
Size : 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[ 6] Habitat : Shrubland and grassland[ 20] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
LC
Unknown [ 20]
Bailey's shrew
C. baileyi Osgood , 1936
Ethiopia
Size : 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[ 6] Habitat : Grassland[ 21] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
LC
Unknown [ 21]
Bale shrew
C. bottegoides Hutterer & Yalden , 1990
Ethiopia
Size : 4–6 cm (2 in) long, plus 4–5 cm (2 in) tail[ 22] Habitat : Grassland and forest[ 23] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
EN
Unknown [ 23]
Banka shrew
C. vosmaeri Jentink , 1888
Bangka Island in Indonesia
Size : 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[ 6] Habitat : Forest[ 24] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
DD
Unknown [ 24]
Batak shrew
C. batakorum Hutterer , 2007
Philippines
Size : 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[ 6] Habitat : Forest[ 25] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
DD
Unknown [ 25]
Bates's shrew
C. batesi (Dollman , 1915)
Western Africa
Size : 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[ 6] Habitat : Forest[ 26] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
LC
Unknown [ 26]
Beccari's shrew
C. beccarii Dobson , 1886
Sumatra
Size : 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[ 6] Habitat : Forest[ 27] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
LC
Unknown [ 27]
Beletta shrew
C. yaldeni Lavrenchenko , Voyta , & Hutterer , 2016
Ethiopia
Size : 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[ 6] Habitat : Forest[ 28] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
DD
Unknown [ 28]
Bicolored musk shrew
C. fuscomurina (Heuglin , 1865)
Sub-Saharan Africa
Size : 4–8 cm (2–3 in) long, plus 2–5 cm (1–2 in) tail[ 29] Habitat : Savanna, grassland, and desert[ 30] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
LC
Unknown [ 30]
Bicolored shrew
C. leucodon (Hermann , 1780)
Europe and western Asia
Size : 6–9 cm (2–4 in) long, plus 2–5 cm (1–2 in) tail[ 31] Habitat : Shrubland and grassland[ 32] Diet : Small mammals, frogs, toads, lizards, and invertebrates[ 31]
LC
Unknown [ 32]
Black-footed shrew
C. nigripes Miller & Hollister , 1921
C. n. lipara
C. n. nigripes
Sulawesi island in Indonesia
Size : 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[ 6] Habitat : ForestDiet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
LC
Unknown
Blackish white-toothed shrew
C. nigricans Bocage , 1889
Angola
Size : 6–9 cm (2–4 in) long, plus 4–7 cm (2–3 in) tail[ 34] Habitat : Unknown[ 35] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
LC
Unknown [ 35]
Bornean shrew
C. foetida Peters , 1870
C. f. doriae
C. f. foetida
C. f. kelabit
Borneo
Size : 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[ 6] Habitat : Forest[ 36] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
LC
Unknown [ 36]
Bottego's shrew
C. bottegi Thomas , 1898
Ethiopia
Size : 4–6 cm (2 in) long, plus 2–3 cm (1 in) tail[ 37] Habitat : Unknown[ 38] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
DD
Unknown [ 38]
Buettikofer's shrew
C. buettikoferi Jentink , 1888
West-central Africa
Size : 7–10 cm (3–4 in) long, plus 5–7 cm (2–3 in) tail[ 39] Habitat : ForestDiet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
LC
Unknown
Butiaba naked-tailed shrew
C. littoralis Heller , 1910
Central Africa
Size : 8–10 cm (3–4 in) long, plus 6–8 cm (2–3 in) tail[ 41] Habitat : Forest[ 42] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
LC
Unknown [ 42]
Cameroonian shrew
C. picea Sanderson , 1940
West-central Africa
Size : 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[ 6] Habitat : Forest and grassland[ 43] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
EN
Unknown [ 43]
Canarian shrew
C. canariensis Hutterer , López-Jurado , & Vogel , 1987
Canary Islands
Size : 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[ 6] Habitat : Coastal marine, desert, and shrubland[ 44] Diet : Snails and insects[ 44]
EN
Unknown [ 44]
Caspian shrew
C. caspica Thomas , 1907
Iran and Azerbaijan
Size : 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[ 6] Habitat : Unknown[ 45] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
DD
Unknown [ 45]
Chinese white-toothed shrew
C. rapax Allen , 1923
C. r. kurodai
C. r. lutaoensis
C. r. rapax
C. r. tadae
Eastern Asia
Size : 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[ 6] Habitat : Unknown[ 46] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
DD
Unknown [ 46]
Christmas Island shrew
C. trichura Dobson , 1889
Christmas Island
Size : 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[ 6] Habitat : Forest[ 47] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
CR
0–200 [ 47]
Cinderella shrew
C. cinderella Thomas , 1911
Western Africa
Size : 5–7 cm (2–3 in) long, plus 4–5 cm (2–2 in) tail[ 48] Habitat : Savanna[ 49] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
LC
Unknown [ 49]
Congo white-toothed shrew
C. congobelgica Hollister , 1916
Central Africa
Size : 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[ 6] Habitat : Forest[ 50] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
LC
Unknown [ 50]
Cranbrook's white-toothed shrew
C. cranbrooki Jenkins , Lunde , & Moncrieff , 2009
Northern Myanmar
Size : 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[ 6] Habitat : Forest[ 51] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
DD
Unknown [ 51]
Cretan shrew
C. zimmermanni Wettstein , 1953
Island of Crete in Greece
Size : 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[ 6] Habitat : Unknown and rocky areas[ 52] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
EN
Unknown [ 52]
Crosse's shrew
C. crossei Thomas , 1895
Western Africa
Size : 6–7 cm (2–3 in) long, plus 4–6 cm (2 in) tail[ 53] Habitat : Forest[ 54] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
LC
Unknown [ 54]
Cyrenaica shrew
C. aleksandrisi Vesmanis , 1977
Northern Libya
Size : 5–7 cm (2–3 in) long, plus 3–5 cm (1–2 in) tail[ 55] Habitat : Shrubland[ 56] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
LC
Unknown [ 56]
Dent's shrew
C. denti Dollman , 1915
Western and central Africa
Size : About 6 cm (2 in) long, plus 5 cm (2 in) tail[ 57] Habitat : Forest and savanna[ 58] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
LC
Unknown [ 58]
Desert musk shrew
C. smithii Thomas , 1895
C. s. debalsaci
C. s. smithii
Western and Eastern Africa
Size : 6–9 cm (2–4 in) long, plus 3–6 cm (1–2 in) tail[ 59] Habitat : Savanna[ 60] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
LC
Unknown [ 60]
Desperate shrew
C. desperata Hutterer , Jenkins , & Verheyen , 1991
Tanzania
Size : 8–11 cm (3–4 in) long, plus about 7 cm (3 in) tail[ 61] Habitat : Forest[ 62] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
VU
Unknown [ 62]
Dhofar shrew
C. dhofarensis (Hutterer , 1988)
Yemen and Oman
Size : 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[ 6] Habitat : Unknown[ 63] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
DD
Unknown [ 63]
Doucet's musk shrew
C. douceti Heim de Balsac , 1958
Western Africa
Size : 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[ 6] Habitat : Forest[ 64] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
LC
Unknown [ 64]
Dsinezumi shrew
C. dsinezumi (Temminck , 1842)
Japan
Size : 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[ 6] Habitat : Shrubland, grassland, and inland wetlands[ 65] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
LC
Unknown [ 65]
East African highland shrew
C. allex Osgood , 1910
Kenya and Tanzania
Size : 5–7 cm (2–3 in) long, plus 4–6 cm (2 in) tail[ 66] Habitat : Grassland and inland wetlands[ 67] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
LC
Unknown [ 67]
Egyptian pygmy shrew
C. religiosa (Geoffroy , 1827)
Northern Egypt
Size : 4–7 cm (2–3 in) long, plus 2–4 cm (1–2 in) tail[ 68] Habitat : [ 69] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
DD
Unknown [ 69]
Eisentraut's shrew
C. eisentrauti Heim de Balsac , 1957
Cameroon
Size : 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[ 6] Habitat : Grassland and forest[ 70] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
VU
Unknown [ 70]
Elgon shrew
C. elgonius Osgood , 1910
Kenya and Tanzania
Size : 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[ 6] Habitat : Forest[ 71] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
LC
Unknown [ 71]
Elongated shrew
C. elongata Miller & Hollister , 1921
Sulawesi island in Indonesia
Size : 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[ 6] Habitat : Forest and shrubland[ 72] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
LC
Unknown [ 72]
Fingui white-toothed shrew
C. fingui Ceríaco et al., 2015
Príncipe in São Tomé and Príncipe
Size : 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[ 6] Habitat : Forest[ 73] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
DD
Unknown [ 73]
Fischer's shrew
C. fischeri Pagenstecher , 1885
Kenya and Tanzania
Size : 6–10 cm (2–4 in) long, plus 4–6 cm (2 in) tail[ 74] Habitat : SavannaDiet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
DD
Unknown
Flat-headed shrew
C. planiceps Heller , 1910
West-central Africa
Size : 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[ 6] Habitat : Inland wetlands[ 76] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
DD
Unknown [ 76]
Flower's shrew
C. floweri Dollman , 1915
Northern Egypt
Size : 5–8 cm (2–3 in) long, plus 5–6 cm (2–2 in) tail[ 77] Habitat : Artificial terrainDiet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
DD
Unknown
Fox's shrew
C. foxi Dollman , 1915
Western and central Africa
Size : 9–12 cm (4–5 in) long, plus 5–8 cm (2–3 in) tail[ 79] Habitat : Savanna and forest[ 80] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
LC
Unknown [ 80]
Fraser's musk shrew
C. poensis (Fraser , 1843)
Western Africa
Size : 7–10 cm (3–4 in) long, plus 4–7 cm (2–3 in) tail[ 81] Habitat : Shrubland and forest[ 82] Diet : Arthropods and other invertebrates[ 81]
LC
Unknown [ 82]
Gathorne's shrew
C. gathornei Jenkins , 2013
Himalayas in India
Size : 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[ 6] Habitat : Shrubland and grassland[ 83] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
DD
Unknown [ 83]
Glass's shrew
C. glassi Heim de Balsac , 1966
Ethiopia
Size : 6–9 cm (2–4 in) long, plus 4–6 cm (2 in)tail[ 84] Habitat : Shrubland, grassland, and inland wetlands[ 85] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
NT
Unknown [ 85]
Gmelin's white-toothed shrew
C. gmelini Pallas , 1811
Central Asia
Size : 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[ 6] Habitat : Shrubland and grassland[ 86] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
LC
Unknown [ 86]
Goliath shrew
C. goliath Thomas , 1906
Central Africa
Size : 15–18 cm (6–7 in) long, plus 10–14 cm (4–6 in) tail[ 87] Habitat : Forest[ 88] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
LC
Unknown [ 88]
Gracile naked-tailed shrew
C. maurisca Thomas , 1904
Central Africa
Size : 7–10 cm (3–4 in) long, plus 6–8 cm (2–3 in) tail[ 89] Habitat : Inland wetlands and forest[ 90] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
LC
Unknown [ 90]
Grasse's shrew
C. grassei Brosset , Dubost , & Heim de Balsac , 1965
West-central Africa
Size : 7–9 cm (3–4 in) long, plus 6–8 cm (2–3 in) tail[ 91] Habitat : Forest[ 92] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
LC
Unknown [ 92]
Greater Mindanao shrew
C. grandis Miller , 1911
Philippines
Size : 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[ 6] Habitat : Forest[ 93] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
DD
Unknown [ 93]
Greater red musk shrew
C. flavescens (Geoffroy , 1827)
Southern Africa
Size : 8–12 cm (3–5 in) long, plus 3–6 cm (1–2 in) tail[ 94] Habitat : Savanna and grassland[ 95] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
LC
Unknown [ 95]
Greater white-toothed shrew
C. russula (Hermann , 1780)
Europe and northwestern Africa
Size : 6–9 cm (2–4 in) long, plus 3–5 cm (1–2 in) tail[ 96] Habitat : Shrubland[ 97] Diet : Invertebrates, as well as small mammals and small lizards[ 96]
LC
Unknown [ 97]
Greenwood's shrew
C. greenwoodi Heim de Balsac , 1966
Somalia
Size : 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[ 6] Habitat : Forest, savanna, and shrubland[ 98] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
LC
Unknown [ 98]
Guramba shrew
C. phaeura Osgood , 1936
Ethiopia
Size : 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[ 6] Habitat : Forest[ 99] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
EN
Unknown [ 99]
Guy's shrew
C. guy Jenkins , Lunde , & Moncrieff , 2009
Vietnam
Size : 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[ 6] Habitat : Forest[ 100] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
DD
Unknown [ 100]
Güldenstädt's shrew
C. gueldenstaedtii (Pallas , 1811)
Europe and western Asia
Size : 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[ 6] Habitat : Shrubland, grassland, inland wetlands, rocky areas, intertidal marine , and coastal marine[ 101] Diet : Insects, as well as molluscs, amphipods , annelids , and small vertebrates[ 101]
LC
Unknown [ 101]
Hainan Island shrew
C. wuchihensis Sung , 1966
Southern China and northern Vietnam
Size : 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[ 6] Habitat : Forest[ 102] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
DD
Unknown [ 102]
Harenna shrew
C. harenna Hutterer & Yalden , 1990
Ethiopia
Size : 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[ 6] Habitat : Forest[ 103] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
CR
Unknown [ 103]
Heather shrew
C. erica Dollman , 1915
Angola
Size : About 10 cm (4 in) long, plus 5 cm (2 in) tail[ 104] Habitat : Unknown[ 105] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
DD
Unknown [ 105]
Hildegarde's shrew
C. hildegardeae Thomas , 1904
Central Africa
Size : 6–8 cm (2–3 in) long, plus 3–6 cm (1–2 in) tail[ 106] Habitat : Forest[ 107] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
LC
Unknown [ 107]
Hill's shrew
C. hilliana Jenkins & Smith , 1995
Thailand and Laos
Size : 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[ 6] Habitat : Forest and shrubland[ 108] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
DD
Unknown [ 108]
Horsfield's shrew
C. horsfieldii (Tomes , 1856)
Scattered southern Asia
Size : 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[ 6] Habitat : Forest[ 109] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
DD
Unknown [ 109]
Hun shrew
C. attila Dollman , 1915
Central Africa
Size : 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[ 6] Habitat : Forest[ 110] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
LC
Unknown [ 110]
Hutan shrew
C. hutanis Ruedi & Vogel , 1995
Sumatra island in Indonesia
Size : 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[ 6] Habitat : Forest[ 111] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
LC
Unknown [ 111]
Indochinese shrew
C. indochinensis Kloss , 1922
Southeastern Asia
Size : 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[ 6] Habitat : Forest[ 112] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
LC
Unknown [ 112]
Iranian shrew
C. susiana Redding & Lay , 1978
Iran
Size : 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[ 6] Habitat : Unknown[ 113] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
DD
Unknown [ 113]
Jackass shrew
C. arispa Spitzenberger , 1971
Turkey
Size : 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[ 6] Habitat : Rocky areas[ 114] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
LC
Unknown [ 114]
Jackson's shrew
C. jacksoni Thomas , 1904
East-central Africa
Size : 5–9 cm (2–4 in) long, plus 4–7 cm (2–3 in) tail[ 115] Habitat : Forest[ 116] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
LC
Unknown [ 116]
Javan ghost shrew
C. umbra Demos , Achmadi , Handika , Maharadatunkamsi , Rowe , & Esselstyn , 2016
Java island in Indonesia
Size : 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[ 6] Habitat : Forest[ 117] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
NT
Unknown [ 117]
Javanese shrew
C. maxi Sody , 1936
Indonesia and East Timor
Size : 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[ 6] Habitat : Forest[ 118] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
LC
Unknown [ 118]
Jenkins's shrew
C. jenkinsi Chakraborty , 1978
South Andaman Island in India
Size : 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[ 6] Habitat : Forest[ 119] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
CR
Unknown [ 119]
Jouvenet's shrew
C. jouvenetae Heim de Balsac , 1958
Western Africa
Size : 6–9 cm (2–4 in) long, plus 4–7 cm (2–3 in) tail[ 120] Habitat : Forest[ 121] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
LC
Unknown [ 121]
Kashmir white-toothed shrew
C. pullata Miller , 1911
Northern India and Pakistan
Size : 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[ 6] Habitat : Unknown[ 122] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
DD
Unknown [ 122]
Katinka's shrew
C. katinka Bate , 1937
Western Asia
Size : 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[ 6] Habitat : Unknown[ 123] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
DD
Unknown [ 123]
Kilimanjaro shrew
C. monax Thomas , 1910
Size : 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[ 6] Habitat : Forest[ 124] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
DD
Unknown [ 124]
Kinabalu shrew
C. baluensis Thomas , 1898
Northern Borneo
Size : 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[ 6] Habitat : Forest, shrubland, and grassland[ 125] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
LC
Unknown [ 125]
Kivu long-haired shrew
C. lanosa Heim de Balsac , 1968
Central Africa
Size : 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[ 6] Habitat : Forest and inland wetlands[ 126] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
VU
Unknown [ 126]
Kivu shrew
C. kivuana Heim de Balsac , 1968
Central Africa
Size : 6–9 cm (2–4 in) long, plus 6–8 cm (2–3 in) tail[ 127] Habitat : Forest and inland wetlands[ 128] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
NT
Unknown [ 128]
Lamotte's shrew
C. lamottei Heim de Balsac , 1968
Western Africa
Size : 8–9 cm (3–4 in) long, plus 3–6 cm (1–2 in) tail[ 127] Habitat : Forest and savanna[ 129] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
LC
Unknown [ 129]
Large-headed forest shrew
C. grandiceps Hutterer , 1983
Western Africa
Size : 9–12 cm (4–5 in) long, plus 5–8 cm (2–3 in) tail[ 130] Habitat : Forest[ 131] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
LC
Unknown [ 131]
Latona's shrew
C. latona Hollister , 1916
Democratic Republic of the Congo
Size : 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[ 6] Habitat : Forest[ 132] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
LC
Unknown [ 132]
Lesser Ryukyu shrew
C. watasei Kuroda , 1924
Southern Japan
Size : 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[ 6] Habitat : Shrubland, grassland, and inland wetlands[ 133] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
LC
Unknown [ 133]
Lesser gray-brown musk shrew
C. silacea Thomas , 1895
Southern Africa
Size : 6–8 cm (2–3 in) long, plus 4–6 cm (2 in) tail[ 134] Habitat : Rocky areas, grassland, shrubland, and savanna[ 135] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
LC
Unknown [ 135]
Lesser red musk shrew
C. hirta Peters , 1852
Southern and southeastern Africa
Size : 6–10 cm (2–4 in) long, plus 3–6 cm (1–2 in) tail[ 136] Habitat : Savanna, grassland, and desert[ 137] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
LC
Unknown [ 137]
Lesser rock shrew
C. serezkyensis Laptev , 1929
Western Asia
Size : 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[ 6] Habitat : Unknown[ 138] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
LC
Unknown [ 138]
Lesser white-toothed shrew
C. suaveolens (Pallas , 1811)
Europe and Asia
Size : 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[ 6] Habitat : Forest, shrubland, grassland, rocky areas, and coastal/supratidal marine [ 139] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
LC
Unknown [ 139]
Long-footed shrew
C. crenata Brosset , Dubost , & Heim de Balsac , 1965
West-central Africa
Size : 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[ 6] Habitat : ForestDiet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
LC
Unknown
Long-tailed musk shrew
C. dolichura Peters , 1876
Central Africa
Size : 6–7 cm (2–3 in) long, plus 7–8 cm (3 in) tail[ 141] Habitat : Forest[ 142] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
LC
Unknown [ 142]
Lucina's shrew
C. lucina Dippenaar , 1980
Ethiopia
Size : 8–10 cm (3–4 in) long, plus 4–6 cm (2 in) tail[ 143] Habitat : Grassland and inland wetlands[ 144] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
VU
Unknown [ 144]
Ludia's shrew
C. ludia Hollister , 1916
Central Africa
Size : 6–7 cm (2–3 in) long, plus 5–6 cm (2–2 in) tail[ 145] Habitat : Forest[ 146] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
LC
Unknown [ 146]
Luzon shrew
C. grayi Dobson , 1890
Philippines
Size : 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[ 6] Habitat : Forest[ 147] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
LC
Unknown [ 147]
Lwiro shrew
C. lwiroensis Peterhans & Hutterer , 2013
Democratic Republic of the Congo
Size : 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[ 6] Habitat : Forest[ 148] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
DD
Unknown [ 148]
MacArthur's shrew
C. macarthuri St. Leger , 1934
Eastern Africa
Size : 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[ 6] Habitat : SavannaDiet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
LC
Unknown
MacMillan's shrew
C. macmillani Dollman , 1915
Ethiopia
Size : 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[ 6] Habitat : Forest and savanna[ 150] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
NT
Unknown [ 150]
Makwassie musk shrew
C. maquassiensis Roberts , 1946
Southern Africa
Size : 5–8 cm (2–3 in) long, plus 4–5 cm (2–2 in) tail[ 151] Habitat : Grassland and rocky areas[ 152] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
LC
Unknown [ 152]
Malayan shrew
C. malayana Robinson & Kloss , 1911
Malaysia
Size : 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[ 6] Habitat : Forest[ 153] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
LC
Unknown [ 153]
Mamfe shrew
C. virgata Sanderson , 1940
West-central Africa
Size : 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[ 6] Habitat : Forest and grassland[ 154] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
LC
Unknown [ 154]
Manenguba shrew
C. manengubae Hutterer , 1982
West-central Africa
Size : 7–9 cm (3–4 in) long, plus 6–7 cm (2–3 in) tail[ 151] Habitat : Forest, shrubland, and grassland[ 155] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
VU
Unknown [ 155]
Mauritanian shrew
C. lusitania Dollman , 1915
Western and eastern Africa
Size : 4–7 cm (2–3 in) long, plus 2–4 cm (1–2 in) tail[ 156] Habitat : Savanna and shrubland[ 157] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
LC
Unknown [ 157]
Mduma's shrew
C. mdumai Stanley , Hutterer , Giarla , & Esselstyn , 2015
Tanzania
Size : 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[ 6] Habitat : Forest[ 158] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
EN
Unknown [ 158]
Mindanao shrew
C. beatus Miller , 1910
Philippines
Size : 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[ 6] Habitat : Forest[ 159] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
LC
Unknown [ 159]
Mindoro shrew
C. mindorus Miller , 1910
Philippines
Size : 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[ 6] Habitat : Forest[ 160] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
DD
Unknown [ 160]
Montane white-toothed shrew
C. montis Thomas , 1906
Eastern Africa
Size : 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[ 6] Habitat : Forest, shrubland, and grassland[ 161] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
LC
Unknown [ 161]
Moonshine shrew
C. luna Dollman , 1910
Southeastern Africa
Size : 6–11 cm (2–4 in) long, plus 3–7 cm (1–3 in) tail[ 156] Habitat : Forest[ 162] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
LC
Unknown [ 162]
Mossy forest shrew
C. musseri Ruedi & Vogel , 1995
Sulawesi island in Indonesia
Size : 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[ 6] Habitat : Forest[ 163] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
DD
Unknown [ 163]
Mount Nimba giant forest shrew
C. nimbasilvanus Hutterer , 2003
Western Africa
Size : 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[ 6] Habitat : Forest[ 164] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
LC
Unknown [ 164]
Munissi's shrew
C. munissii Stanley , Hutterer , Giarla , & Esselstyn , 2015
Tanzania
Size : 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[ 6] Habitat : Forest[ 165] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
LC
Unknown [ 165]
Narrow-headed shrew
C. stenocephala Heim de Balsac & Dieterlen , 1979
Central Africa
Size : 8–11 cm (3–4 in) long, plus 6–9 cm (2–4 in) tail[ 166] Habitat : Inland wetlands[ 167] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
VU
Unknown [ 167]
Negev shrew
C. ramona Ivanitskaya , Shenbrot , & Nevo , 1996
Israel
Size : 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[ 6] Habitat : Desert[ 168] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
LC
Unknown [ 168]
Negros shrew
C. negrina Rabor , 1952
Philippines
Size : 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[ 6] Habitat : Forest[ 169] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
EN
Unknown [ 169]
Nicobar shrew
C. nicobarica Miller , 1902
Great Nicobar Island in India
Size : 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[ 6] Habitat : Forest[ 170] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
CR
Unknown [ 170]
Nigerian shrew
C. nigeriae Dollman , 1915
West-central Africa
Size : 9–11 cm (4 in) long, plus 5–7 cm (2–3 in) tail[ 171] Habitat : Forest[ 172] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
LC
Unknown [ 172]
Nimba shrew
C. nimbae Heim de Balsac , 1956
Western Africa
Size : 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[ 6] Habitat : Forest[ 173] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
NT
Unknown [ 173]
Niobe's shrew
C. niobe Thomas , 1906
Central Africa
Size : 6–9 cm (2–4 in) long, plus 5–7 cm (2–3 in) tail[ 174] Habitat : Forest and grassland[ 175] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
LC
Unknown [ 175]
North African white-toothed shrew
C. pachyura Küster , 1835
Northern Africa and southern Europe
Size : 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[ 6] Habitat : Shrubland[ 176] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
LC
Unknown [ 176]
Nyiro shrew
C. macowi Dollman , 1915
Kenya
Size : 6–8 cm (2–3 in) long, plus 5–6 cm (2–2 in) tail[ 177] Habitat : Unknown[ 178] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
DD
Unknown [ 178]
Oriental shrew
C. orientalis Jentink , 1890
C. o. lawuana
C. o. orientalis
Indonesia
Size : 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[ 6] Habitat : Forest[ 179] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
LC
Unknown [ 179]
Palawan shrew
C. palawanensis Taylor , 1934
Philippines
Size : 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[ 6] Habitat : Forest and shrubland[ 180] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
LC
Unknown [ 180]
Pale gray shrew
C. pergrisea Miller , 1913
Pakistan
Size : 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[ 6] Habitat : Forest[ 181] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
DD
Unknown [ 181]
Panay shrew
C. panayensis Hutterer , 2007
Philippines
Size : 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[ 6] Habitat : Forest[ 182] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
DD
Unknown [ 182]
Peninsular shrew
C. negligens Kloss , 1914
Southeastern Asia
Size : 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[ 6] Habitat : Forest[ 183] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
LC
Unknown [ 183]
Peters's musk shrew
C. gracilipes Peters , 1870
Tanzania
Size : About 6 cm (2 in) long, plus 5 cm (2 in) tail[ 87] Habitat : UnknownDiet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
DD
Unknown
Phan Luong's shrew
C. phanluongi Jenkins , Abramov , Rozhnov , & Olsson , 2010
Vietnam and Cambodia
Size : 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[ 6] Habitat : Forest[ 185] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
LC
Unknown [ 185]
Phu Hoc shrew
C. phuquocensis Abramov , Jenkins , Rozhnov , & Kalinin , 2008
Phú Quốc island in Vietnam
Size : 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[ 6] Habitat : Forest[ 186] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
VU
Unknown [ 186]
Pitman's shrew
C. pitmani Barclay, 1932
Zambia
Size : About 5 cm (2 in) long, plus 4 cm (2 in) tail[ 187] Habitat : Unknown[ 188] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
DD
Unknown [ 188]
Polia's shrew
C. polia Hollister , 1916
Democratic Republic of the Congo
Size : About 6 cm (2 in) long, plus 7 cm (3 in) tail[ 81] Habitat : Unknown[ 189] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
DD
Unknown [ 189]
Rainey's shrew
C. raineyi Heller , 1912
Kenya
Size : 9–10 cm (4 in) long, plus 5–7 cm (2–3 in) tail[ 190] Habitat : Forest[ 191] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
DD
Unknown [ 191]
Reddish-gray musk shrew
C. cyanea (Duvernoy , 1838)
Southern Africa
Size : 6–10 cm (2–4 in) long, plus 3–7 cm (1–3 in) tail[ 192] Habitat : Forest, grassland, and caves[ 193] Diet : Invertebrates[ 193]
LC
Unknown [ 193]
Roosevelt's shrew
C. roosevelti (Heller , 1910)
Central Africa
Size : 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[ 6] Habitat : Savanna[ 194] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
LC
Unknown [ 194]
Ryukyu shrew
C. orii Kuroda , 1924
Southern Japan
Size : 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[ 6] Habitat : Forest[ 195] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
EN
Unknown [ 195]
Sa Pa shrew
C. sapaensis Jenkins , Abramov , Bannikova , & Rozhnov , 2013
Vietnam
Size : 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[ 6] Habitat : Forest and grassland[ 196] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
DD
Unknown [ 196]
Saharan shrew
C. tarfayensis Vesmanis & Vesmanis , 1980
Mauritania and Morocco
Size : 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[ 6] Habitat : Rocky areas and desert[ 197] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
DD
Unknown [ 197]
Sahelian tiny shrew
C. pasha Dollman , 1915
Scattered northern Africa
Size : 4–6 cm (2 in) long, plus 2–4 cm (1–2 in) tail[ 198] Habitat : Savanna[ 199] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
LC
Unknown [ 199]
Savanna dwarf shrew
C. nanilla Thomas , 1909
Western and eastern Africa
Size : 4–6 cm (2–2 in) long, plus 3–5 cm (1–2 in) tail[ 200] Habitat : Savanna[ 201] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
LC
Unknown [ 201]
Savanna path shrew
C. viaria (Geoffroy , 1834)
Western, central, and eastern Africa
Size : 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[ 6] Habitat : Forest and savanna[ 202] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
LC
Unknown [ 202]
Savanna shrew
C. fulvastra (Sundevall , 1843)
Central Africa
Size : 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[ 6] Habitat : Savanna[ 203] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
LC
Unknown [ 203]
Savanna swamp shrew
C. longipes Hutterer & Happold , 1983
West-central Africa
Size : 9–12 cm (4–5 in) long, plus 5–7 cm (2–3 in) tail[ 204] Habitat : Inland wetlands[ 205] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
DD
Unknown [ 205]
Siberian shrew
C. sibirica Dukelsky , 1930
Central Asia
Size : 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[ 6] Habitat : Forest, grassland, inland wetlands, and desert[ 206] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
LC
Unknown [ 206]
Sibuyan shrew
C. ninoyi Esselstyn & Goodman , 2010
Philippines
Size : 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[ 6] Habitat : Forest[ 207] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
DD
Unknown [ 207]
Sicilian shrew
C. sicula Miller , 1900
C. s. aegatensis
C. s. calypso
C. s. esuae
C. s. sicula
Italy and Malta
Size : 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[ 6] Habitat : Shrubland[ 208] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
LC
Unknown [ 208]
Sinharaja white-toothed shrew
C. hikmiya Meegaskumbura , Meegaskumbura , Pethiyagoda , Manamendra-Arachchi , & Schneider , 2007
Sri Lanka
Size : 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[ 6] Habitat : Forest[ 209] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
EN
Unknown [ 209]
Small-footed shrew
C. parvipes Osgood , 1910
Central Africa
Size : 6–9 cm (2–4 in) long, plus 3–6 cm (1–2 in) tail[ 210] Habitat : Forest, savanna, and shrublandDiet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
LC
Unknown
Smoky white-toothed shrew
C. fumosa Thomas , 1904
Kenya
Size : 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[ 6] Habitat : Forest[ 212] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
LC
Unknown [ 212]
Sokolov's shrew
C. sokolovi Jenkins , Abramov , Rozhnov , & Makarova , 2007
Vietnam
Size : 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[ 6] Habitat : Forest[ 213] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
NT
Unknown [ 213]
Somali dwarf shrew
C. nana Dobson , 1890
Ethiopia and Somalia
Size : About 4 cm (2 in) long, plus 3 cm (1 in) tail[ 214] Habitat : Grassland[ 215] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
DD
Unknown [ 215]
Somali shrew
C. somalica Thomas , 1895
Eastern Africa
Size : 6–8 cm (2–3 in) long, plus 4–5 cm (2–2 in) tail[ 166] Habitat : Savanna[ 216] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
LC
Unknown [ 216]
Southeast Asian shrew
C. fuliginosa (Blyth , 1856)
Southeastern Asian
Size : 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[ 6] Habitat : Forest, shrubland, grassland, and rocky areas[ 217] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
LC
Unknown [ 217]
Sri Lankan long-tailed shrew
C. miya Phillips , 1929
Sri Lanka
Size : 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[ 6] Habitat : Forest[ 218] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
EN
Unknown [ 218]
Sulawesi shrew
C. lea Miller & Hollister , 1921
Sulawesi island in Indonesia
Size : 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[ 6] Habitat : Forest[ 219] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
LC
Unknown [ 219]
Sulawesi tiny shrew
C. levicula Miller & Hollister , 1921
Sulawesi island in Indonesia
Size : 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[ 6] Habitat : Forest[ 220] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
LC
Unknown [ 220]
Sulawesi white-handed shrew
C. rhoditis Miller & Hollister , 1921
Sulawesi island in Indonesia
Size : 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[ 6] Habitat : Forest[ 221] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
LC
Unknown [ 221]
Sumatran giant shrew
C. lepidura Lyon , 1908
Sumatra island in Indonesia
Size : 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[ 6] Habitat : Forest and shrubland[ 222] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
LC
Unknown [ 222]
Sumatran long-tailed shrew
C. paradoxura Dobson , 1886
Sumatra island in Indonesia
Size : 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[ 6] Habitat : Forest[ 223] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
LC
Unknown [ 223]
Sunda shrew
C. monticola Peters , 1870
Southeastern Asia
Size : 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[ 6] Habitat : Forest[ 224] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
LC
Unknown [ 224]
Swamp musk shrew
C. mariquensis (Smith , 1844)
C. m. mariquensis
C. m. neavei
C. m. shortridgei
Southern Africa
Size : 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[ 6] Habitat : Inland wetlands[ 225] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
LC
Unknown [ 225]
São Tomé shrew
C. thomensis (Bocage , 1887)
São Tomé Island in São Tomé and Príncipe
Size : 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[ 6] Habitat : Forest[ 226] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
EN
Unknown [ 226]
Taiwanese gray shrew
C. tanakae Kuroda , 1938
Eastern Asia
Size : 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[ 6] Habitat : Forest and grassland[ 227] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
LC
Unknown [ 227]
Tanzanian shrew
C. tansaniana Hutterer , 1986
Tanzania
Size : 8–11 cm (3–4 in) long, plus 5–8 cm (2–3 in) tail[ 228] Habitat : Forest[ 229] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
NT
Unknown [ 229]
Tarella shrew
C. tarella Dollman , 1915
Central Africa
Size : 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[ 6] Habitat : Forest[ 230] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
EN
Unknown [ 230]
Telford's shrew
C. telfordi Hutterer , 1986
Tanzania
Size : 10–11 cm (4 in) long, plus 7–9 cm (3–4 in) tail[ 231] Habitat : Forest[ 232] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
VU
Unknown [ 232]
Thalia's shrew
C. thalia Dippenaar , 1980
Ethiopia
Size : 7–10 cm (3–4 in) long, plus 5–7 cm (2–3 in) tail[ 231] Habitat : Forest, savanna, shrubland, grassland, and inland wetlands[ 233] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
LC
Unknown [ 233]
Therese's shrew
C. theresae Heim de Balsac , 1968
Western Africa
Size : 7–11 cm (3–4 in) long, plus 3–6 cm (1–2 in) tail[ 234] Habitat : Forest and savanna[ 235] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
LC
Unknown [ 235]
Thick-tailed shrew
C. brunnea Jentink , 1888
C. b. brunnea
C. b. pudjonica
Indonesia
Size : 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[ 6] Habitat : Forest[ 236] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
LC
Unknown [ 236]
Timor shrew
C. tenuis (Müller , 1840)
Timor
Size : 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[ 6] Habitat : Forest[ 237] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
DD
Unknown [ 237]
Turbo shrew
C. turba Dollman , 1910
Central Africa
Size : 9–11 cm (4 in) long, plus 5–6 cm (2–2 in) tail[ 238] Habitat : Forest[ 239] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
LC
Unknown [ 239]
Ugandan lowland shrew
C. selina (Dollman , 1915)
Kenya and Uganda
Size : 7–9 cm (3–4 in) long, plus 5–7 cm (2–3 in) tail[ 240] Habitat : Forest and inland wetlands[ 241] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
DD
Unknown [ 241]
Ugandan musk shrew
C. mutesae Heller , 1910
Central Africa
Size : About 11 cm (4 in) long, plus 6 cm (2 in) tail[ 214] Habitat : Forest[ 242] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
DD
Unknown [ 242]
Ultimate shrew
C. ultima Dollman , 1915
Kenya
Size : About 9 cm (4 in) long, plus 6 cm (2 in) tail[ 243] Habitat : Forest[ 244] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
DD
Unknown [ 244]
Upemba shrew
C. zimmeri Osgood , 1936
Democratic Republic of the Congo
Size : 11–13 cm (4–5 in) long, plus 4–6 cm (2–2 in) tail[ 245] Habitat : Unknown[ 246] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
DD
Unknown [ 246]
Usambara shrew
C. usambarae Dippenaar , 1980
Tanzania
Size : 7–9 cm (3–4 in) long, plus 5–7 cm (2–3 in) tail[ 247] Habitat : Forest[ 248] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
VU
Unknown [ 248]
Ussuri white-toothed shrew
C. lasiura (Dobson , 1890)
Eastern Asia
Size : 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[ 6] Habitat : Grassland[ 249] Diet : Insects and other invertebrates, as well as small vertebrates[ 249]
LC
Unknown [ 249]
Voi shrew
C. voi Osgood , 1910
Central and eastern Africa
Size : 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[ 6] Habitat : Savanna[ 250] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
LC
Unknown [ 250]
Voracious shrew
C. vorax Allen , 1923
Southeastern Asia
Size : 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[ 6] Habitat : Forest[ 251] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
LC
Unknown [ 251]
West African long-tailed shrew
C. muricauda (Miller , 1900)
Western Africa
Size : 5–7 cm (2–3 in) long, plus tail[ 252] Habitat : Forest[ 253] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
LC
Unknown [ 253]
West African pygmy shrew
C. obscurior Heim de Balsac , 1958
Western Africa
Size : 4–5 cm (2–2 in) long, plus 3–4 cm (1–2 in) tail[ 254] Habitat : Forest[ 255] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
LC
Unknown [ 255]
Whitaker's shrew
C. whitakeri De Winton , 1898
Northwestern Africa
Size : 5–7 cm (2–3 in) long, plus 2–4 cm (1–2 in) tail[ 256] Habitat : Coastal marine, intertidal marine, and shrublandDiet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
LC
Unknown
Wimmer's shrew
C. wimmeri Heim de Balsac & Aellen , 1958
Ivory Coast
Size : 8–9 cm (3–4 in) long, plus 5–6 cm (2–2 in) tail[ 258] Habitat : ForestDiet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
CR
Unknown
Xanthippe's shrew
C. xantippe Osgood , 1910
Kenya and Tanzania
Size : 8–10 cm (3–4 in) long, plus 5–7 cm (2–3 in) tail[ 260] Habitat : Savanna and shrubland[ 261] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
LC
Unknown [ 261]
Yankari shrew
C. yankariensis Hutterer & Jenkins , 1980
Central and eastern Africa
Size : 5–7 cm (2–3 in) long, plus 3–5 cm (1–2 in) tail[ 260] Habitat : Savanna[ 262] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
LC
Unknown [ 262]
Zaphir's shrew
C. zaphiri Dollman , 1915
Ethiopia and Kenya
Size : About 10 cm (4 in) long, plus 6 cm (2 in) tail[ 263] Habitat : UnknownDiet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
DD
Unknown
Zarudny's rock shrew
C. zarudnyi Ognew , 1928
Central Asia
Size : 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[ 6] Habitat : Desert[ 265] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
LC
Unknown [ 265]
Genus Diplomesodon – Brandt , 1852 – one species
Common name
Scientific name and subspecies
Range
Size and ecology
IUCN status and estimated population
Piebald shrew
D. pulchellum (Lichtenstein , 1823)
Central Asia
Size : 5–8 cm (2–3 in) long, plus 2–4 cm (1–2 in) tail[ 266] Habitat : Desert[ 267] Diet : Insects, as well as small lizards[ 267]
LC
Unknown [ 267]
Genus Feroculus – Kelaart , 1852 – one species
Common name
Scientific name and subspecies
Range
Size and ecology
IUCN status and estimated population
Kelaart's long-clawed shrew
F. feroculus (Kelaart , 1850)
Southern India and Sri Lanka
Size : 10–12 cm (4–5 in) long, plus 5–8 cm (2–3 in) tail[ 268] Habitat : Forest, grassland, and inland wetlands[ 269] Diet : Earthworms, small vertebrates, and plants[ 268]
EN
Unknown [ 269]
Genus Ruwenzorisorex – Hutterer , 1986 – one species
Common name
Scientific name and subspecies
Range
Size and ecology
IUCN status and estimated population
Ruwenzori shrew
R. suncoides (Osgood , 1936)
Central Africa
Size : 9–10 cm (4 in) long, plus 5–7 cm (2–3 in) tail[ 274] Habitat : Inland wetlands and forest[ 275] Diet : Mollusks and small vertebrates[ 268]
VU
Unknown [ 275]
Genus Solisorex – Thomas , 1924 – one species
Common name
Scientific name and subspecies
Range
Size and ecology
IUCN status and estimated population
Pearson's long-clawed shrew
S. pearsoni Thomas , 1924
Sri Lanka
Size : 12–15 cm (5–6 in) long, plus 5–7 cm (2–3 in) tail[ 266] Habitat : Forest and grassland[ 280] Diet : Insects, earthworms, and small vertebrates[ 266]
EN
Unknown [ 280]
Genus Suncus – Ehrenberg , 1832 – eighteen species
Common name
Scientific name and subspecies
Range
Size and ecology
IUCN status and estimated population
Anderson's shrew
S. stoliczkanus (Anderson , 1877)
Southern Asia
Size : 5–9 cm (2–4 in) long, plus 3–10 cm (1–4 in) tail[ 281] Habitat : Forest and grassland[ 282] Diet : Insects[ 281]
LC
Unknown [ 282]
Asian highland shrew
S. montanus (Kelaart , 1850)
Southern India and Sri Lanka
Size : 5–9 cm (2–4 in) long, plus 3–10 cm (1–4 in) tail[ 281] Habitat : Forest and grassland[ 283] Diet : Insects[ 281]
LC
Unknown [ 283]
Asian house shrew
S. murinus (Linnaeus , 1766)
Asia and Africa (introduced in red)
Size : 10–5 cm (4–2 in) long, plus 5–9 cm (2–4 in) tail[ 284] Habitat : Forest, savanna, shrubland, grassland, and inland wetlands[ 285] Diet : Insects and small mammals, as well as plants[ 286]
LC
Unknown [ 285]
Black shrew
S. ater (Medway , 1965)
Northern Borneo
Size : 5–9 cm (2–4 in) long, plus 3–10 cm (1–4 in) tail[ 281] Habitat : Forest[ 287] Diet : Insects[ 281]
DD
Unknown [ 287]
Bornean pygmy shrew
S. hosei (Thomas , 1893)
Borneo
Size : 5–9 cm (2–4 in) long, plus 3–10 cm (1–4 in) tail[ 281] Habitat : Forest[ 288] Diet : Insects[ 281]
DD
Unknown [ 288]
Climbing shrew
S. megalura (Jentink , 1888)
Sub-Saharan Africa
Size : 5–6 cm (2 in) long, plus 8–10 cm (3–4 in) tail[ 284] Habitat : Forest and savanna[ 289] Diet : Insects[ 284]
LC
Unknown [ 289]
Day's shrew
S. dayi (Dodson , 1888)
Southern India
Size : 5–9 cm (2–4 in) long, plus 3–10 cm (1–4 in) tail[ 281] Habitat : Forest and grassland[ 290] Diet : Insects[ 281]
EN
Unknown [ 290]
Etruscan shrew
S. etruscus (Savi , 1822)
Europe, northern Africa, and Asia
Size : 3–5 cm (1–2 in) long, plus 2–3 cm (1 in) tail[ 1] Habitat : Forest, shrubland, grassland, and desert[ 291] Diet : Insects[ 292]
LC
Unknown [ 291]
Flores shrew
S. mertensi Kock , 1974
Flores island in Indonesia
Size : 5–9 cm (2–4 in) long, plus 3–10 cm (1–4 in) tail[ 281] Habitat : Forest[ 293] Diet : Insects[ 281]
EN
Unknown [ 293]
Greater dwarf shrew
S. lixus (Thomas , 1898)
Southern Africa
Size : 5–9 cm (2–4 in) long, plus 3–6 cm (1–2 in) tail[ 281] Habitat : Forest, savanna, shrubland, and grassland[ 294] Diet : Insects[ 281]
LC
Unknown [ 294]
Hutu-Tutsi dwarf shrew
S. hututsi Peterhans & Hutterer , 2009
Central Africa
Size : 4–6 cm (2 in) long, plus 2–3 cm (1 in) tail[ 281] Habitat : Forest and shrubland[ 295] Diet : Insects[ 281]
DD
Unknown [ 295]
Jungle shrew
S. zeylanicus Phillips , 1928
Sri Lanka
Size : 5–9 cm (2–4 in) long, plus 3–10 cm (1–4 in) tail[ 281] Habitat : Forest[ 296] Diet : Insects[ 281]
EN
Unknown [ 296]
Least dwarf shrew
S. infinitesimus (Heller , 1912)
South Africa and Kenya
Size : 5–7 cm (2–3 in) long, plus 2–4 cm (1–2 in) tail[ 281] Habitat : Forest and grassland[ 297] Diet : Insects[ 281]
LC
Unknown [ 297]
Lesser dwarf shrew
S. varilla (Thomas , 1895)
Southern Africa
Size : 5–7 cm (2–3 in) long, plus 2–5 cm (1–2 in) tail[ 281] Habitat : Forest, savanna, shrubland, and grassland[ 298] Diet : Insects[ 281]
LC
Unknown [ 298]
Malayan pygmy shrew
S. malayanus (Kloss , 1917)
Malaysia and Thailand
Size : 5–9 cm (2–4 in) long, plus 3–10 cm (1–4 in) tail[ 281] Habitat : Forest[ 299] Diet : Insects[ 281]
LC
Unknown [ 299]
Remy's pygmy shrew
S. remyi Brosset , Dubost , & Heim de Balsac , 1965
West-central Africa
Size : 4–5 cm (2 in) long, plus 1–3 cm (0.4–1.2 in) tail[ 281] Habitat : Forest[ 300] Diet : Insects[ 281]
LC
Unknown [ 300]
Sri Lankan shrew
S. fellowesgordoni Phillips , 1932
Sri Lanka
Size : 5–9 cm (2–4 in) long, plus 3–10 cm (1–4 in) tail[ 281] Habitat : Forest and grassland[ 301] Diet : Insects[ 281]
EN
Unknown [ 301]
Taita shrew
S. aequatorius (Heller , 1912)
Kenya
Size : 6–9 cm (2–4 in) long, plus 5–7 cm (2–3 in) tail[ 302] Habitat : Forest[ 303] Diet : Insects[ 304]
EN
Unknown [ 303]
Genus Sylvisorex – Thomas , 1904 – fifteen species
Common name
Scientific name and subspecies
Range
Size and ecology
IUCN status and estimated population
Bamenda pygmy shrew
S. silvanorum (Hutterer , Riegert , & Sedláček , 2009)
Cameroon
Size : 4–10 cm (2–4 in) long, plus 4–9 cm (2–4 in) tail[ 274] Habitat : Forest and grassland[ 305] Diet : Insects[ 306]
DD
Unknown [ 305]
Bioko forest shrew
S. isabellae Heim de Balsac , 1968
West-central Africa
Size : 4–10 cm (2–4 in) long, plus 4–9 cm (2–4 in) tail[ 274] Habitat : Forest and grassland[ 307] Diet : Insects[ 306]
VU
Unknown [ 307]
Cameroonian forest shrew
S. camerunensis Heim de Balsac , 1968
West-central Africa
Size : 5–6 cm (2 in) long, plus 5–7 cm (2–3 in) tail[ 308] Habitat : Forest and grassland[ 309] Diet : Insects[ 306]
VU
Unknown [ 309]
Corbet's forest shrew
S. corbeti Hutterer , 2009
West-central Africa
Size : 4–10 cm (2–4 in) long, plus 4–9 cm (2–4 in) tail[ 274] Habitat : Forest[ 310] Diet : Insects[ 306]
DD
Unknown [ 310]
Dudu Akaibe's pygmy shrew
S. akaibei Mukinzi , Hutterer , & Barriere , 2009
Democratic Republic of the Congo
Size : 4–10 cm (2–4 in) long, plus 4–9 cm (2–4 in) tail[ 274] Habitat : Forest[ 311] Diet : Insects[ 306]
DD
Unknown [ 311]
Grant's forest shrew
S. granti Thomas , 1907
S. g. granti
S. g. mundus
Central Africa
Size : 4–7 cm (2–3 in) long, plus 4–7 cm (2–3 in) tail[ 312] Habitat : Forest[ 313] Diet : Insects[ 306]
LC
Unknown [ 313]
Greater forest shrew
S. ollula Thomas , 1913
Central Africa
Size : 5–8 cm (2–3 in) long, plus 1–2 cm (0–1 in) tail[ 314] Habitat : Forest[ 315] Diet : Insects[ 306]
LC
Unknown [ 315]
Howell's forest shrew
S. howelli Jenkins , 1984
Tanzania
Size : 5–8 cm (2–3 in) long, plus 3–5 cm (1–2 in) tail[ 316] Habitat : Forest[ 317] Diet : Insects[ 306]
LC
Unknown [ 317]
Johnston's forest shrew
S. johnstoni (Dobson , 1888)
West-central and central Africa
Size : 7–10 cm (3–4 in) long, plus 4–7 cm (2–3 in) tail[ 308] Habitat : Forest[ 318] Diet : Insects[ 306]
LC
Unknown [ 318]
Kongana shrew
S. konganensis Ray & Hutterer , 1996
Central Africa
Size : 4–10 cm (2–4 in) long, plus 4–9 cm (2–4 in) tail[ 274] Habitat : Forest[ 319] Diet : Insects[ 306]
DD
Unknown [ 319]
Lesser forest shrew
S. oriundus Hollister , 1916
Central Africa
Size : 4–10 cm (2–4 in) long, plus 4–9 cm (2–4 in) tail[ 274] Habitat : Forest[ 320] Diet : Insects[ 306]
DD
Unknown [ 320]
Moon forest shrew
S. lunaris Thomas , 1906
Central Africa
Size : 7–10 cm (3–4 in) long, plus 4–6 cm (2 in) tail[ 321] Habitat : Forest[ 322] Diet : Insects[ 306]
NT
Unknown [ 322]
Mount Cameroon forest shrew
S. morio (Gray , 1862)
West-central Africa
Size : 6–8 cm (2–3 in) long, plus 5–6 cm (2 in) tail[ 323] Habitat : Forest[ 324] Diet : Insects[ 306]
EN
Unknown [ 324]
Rain forest shrew
S. pluvialis Hutterer & Schlitter , 1996
West-central Africa
Size : 6–8 cm (2–3 in) long, plus 5–8 cm (2–3 in) tail[ 321] Habitat : Forest[ 325] Diet : Insects[ 306]
DD
Unknown [ 325]
Volcano shrew
S. vulcanorum Hutterer & Verheyan , 1985
Central Africa
Size : 4–6 cm (2 in) long, plus 4–7 cm (2–3 in) tail[ 308] Habitat : Forest and inland wetlands[ 326] Diet : Insects[ 306]
LC
Unknown [ 326]
References
^ a b Nowak , p. 224
^ Jenkins, P. D.; Lunde, D. P.; Moncrieff, C. B. (2009). "Descriptions of New Species of Crocidura (Soricomorpha: Soricidae) from Mainland Southeast Asia, with Synopses of Previously Described Species and Remarks on Biogeography" (PDF) . Bulletin of the American Museum of Natural History . 331 : 356–405. doi :10.1206/582-10.1 . S2CID 84803490 . Archived (PDF) from the original on 2020-03-13. Retrieved 2024-04-12 .
^ "Fossilworks: Crocidurinae" . Paleobiology Database . University of Wisconsin–Madison . Archived from the original on June 6, 2023. Retrieved April 11, 2024 .
^ Nations, J. A.; Giarla, T. C.; Morni, M. A.; William Dee, J.; Swanson, M. T.; Hiller, A. E.; Khan, F. A. A.; Esselstyn, J. A. (2022). "Molecular data from the holotype of the enigmatic Bornean Black Shrew, Suncus ater Medway, 1965 (Soricidae, Crocidurinae), place it in the genus Palawanosorex " . ZooKeys (1137): 17–31. Bibcode :2022ZooK.1137...17N . doi :10.3897/zookeys.1137.94217 . PMC 9836656 . PMID 36760481 .
^ Wilson, Reeder , pp. 224-262
^ a b c d e f g h i j k l m n o p q r s t u v w x y z aa ab ac ad ae af ag ah ai aj ak al am an ao ap aq ar as at au av aw ax ay az ba bb bc bd be bf bg bh bi bj bk bl bm bn bo bp bq br bs bt bu bv bw bx by bz ca cb cc cd ce cf cg ch ci cj ck cl cm cn co cp cq cr cs ct cu cv cw cx cy cz da db dc dd de df dg dh di dj dk dl dm dn do dp dq dr ds dt du dv dw dx dy dz ea eb ec ed ee ef eg eh ei ej ek el em en eo ep eq er es et eu ev ew ex ey ez fa fb fc fd fe ff fg fh fi fj fk fl fm fn fo fp fq fr fs ft fu fv fw fx fy fz ga gb gc gd ge gf gg gh gi gj gk gl gm gn go gp gq gr gs gt gu gv gw gx gy gz ha hb hc hd he hf hg hh hi hj hk hl hm hn ho hp hq hr hs ht hu hv hw hx hy hz ia ib ic id ie if ig ih ii ij ik il im in io ip iq ir is it iu iv iw ix iy iz ja jb jc jd je jf jg jh ji jj jk jl jm jn jo jp jq jr js jt ju jv jw jx jy jz ka kb kc kd ke kf kg kh ki kj kk kl km kn Nowak , p. 221
^ a b Cassola, F. (2017) [errata version of 2016 assessment]. "Crocidura nigrofusca " . IUCN Red List of Threatened Species . 2016 : e.T41346A115180069. doi :10.2305/IUCN.UK.2016-3.RLTS.T41346A22310000.en .
^ Kingdon , p. 64
^ a b Hutterer, R. (2016). "Crocidura caliginea " . IUCN Red List of Threatened Species . 2016 : e.T5623A22300539. doi :10.2305/IUCN.UK.2016-2.RLTS.T5623A22300539.en .
^ a b Kingdon , p. 119
^ a b Cassola, F. (2017) [errata version of 2016 assessment]. "Crocidura olivieri " . IUCN Red List of Threatened Species . 2016 : e.T41348A115180235. doi :10.2305/IUCN.UK.2016-3.RLTS.T41348A22309617.en .
^ a b Molur, S. (2017) [errata version of 2016 assessment]. "Crocidura andamanensis " . IUCN Red List of Threatened Species . 2016 : e.T5595A115076487. doi :10.2305/IUCN.UK.2016-3.RLTS.T5595A22303530.en .
^ a b Molur, S. (2017) [errata version of 2016 assessment]. "Crocidura hispida " . IUCN Red List of Threatened Species . 2016 : e.T5603A115076898. doi :10.2305/IUCN.UK.2016-3.RLTS.T5603A22299245.en .
^ a b Dando, T.; Kennerley, R.; Jenkins, P. (2019). "Crocidura annamitensis " . IUCN Red List of Threatened Species . 2019 : e.T48296764A48296771. doi :10.2305/IUCN.UK.2019-1.RLTS.T48296764A48296771.en .
^ Kingdon , p. 58
^ a b Kennerley, R. (2017) [errata version of 2016 assessment]. "Crocidura ansellorum " . IUCN Red List of Threatened Species . 2016 : e.T5558A115073943. doi :10.2305/IUCN.UK.2016-3.RLTS.T5558A22305108.en .
^ a b Amori, G. (2019). "Crocidura arabica " . IUCN Red List of Threatened Species . 2019 : e.T40618A22295560. doi :10.2305/IUCN.UK.2019-1.RLTS.T40618A22295560.en .
^ a b Gerrie, R.; Kennerley, R. (2017) [errata version of 2016 assessment]. "Crocidura armenica " . IUCN Red List of Threatened Species . 2016 : e.T5596A115076665. doi :10.2305/IUCN.UK.2016-3.RLTS.T5596A22303457.en .
^ a b Heaney, L.; Molur, S. (2018) [errata version of 2017 assessment]. "Crocidura attenuata " . IUCN Red List of Threatened Species . 2017 : e.T48296412A123807388. doi :10.2305/IUCN.UK.2017-3.RLTS.T48296412A22295645.en .
^ a b Temple, H. (2017) [errata version of 2016 assessment]. "Crocidura shantungensis " . IUCN Red List of Threatened Species . 2016 : e.T5617A115077696. doi :10.2305/IUCN.UK.2016-3.RLTS.T5617A22300721.en .
^ a b Kennerley, R.; Lavrenchenko, L. (2017) [errata version of 2016 assessment]. "Crocidura baileyi " . IUCN Red List of Threatened Species . 2016 : e.T5559A115074097. doi :10.2305/IUCN.UK.2016-3.RLTS.T5559A22305001.en .
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^ a b Lavrenchenko, L. (2017) [errata version of 2016 assessment]. "Crocidura bottegoides " . IUCN Red List of Threatened Species . 2016 : e.T5622A115078080. doi :10.2305/IUCN.UK.2016-3.RLTS.T5622A22300328.en .
^ a b Gerrie, R.; Kennerley, R. (2017) [errata version of 2016 assessment]. "Crocidura vosmaeri " . IUCN Red List of Threatened Species . 2016 : e.T136447A115207832. doi :10.2305/IUCN.UK.2016-3.RLTS.T136447A22298267.en .
^ a b Dando, T.; Kennerley, R. (2020) [amended version of 2019 assessment]. "Crocidura batakorum " . IUCN Red List of Threatened Species . 2020 : e.T48297939A166612831. doi :10.2305/IUCN.UK.2020-1.RLTS.T48297939A166612831.en .
^ a b Hutterer, R. (2016). "Crocidura batesi " . IUCN Red List of Threatened Species . 2016 : e.T40620A22295796. doi :10.2305/IUCN.UK.2016-2.RLTS.T40620A22295796.en .
^ a b Cassola, F. (2016). "Crocidura beccarii " . IUCN Red List of Threatened Species . 2016 : e.T5598A22302844. doi :10.2305/IUCN.UK.2016-2.RLTS.T5598A22302844.en .
^ a b Dando, T.; Kennerley, R. (2019). "Crocidura yaldeni " . IUCN Red List of Threatened Species . 2019 : e.T112517512A112517515. doi :10.2305/IUCN.UK.2019-1.RLTS.T112517512A112517515.en .
^ Kingdon , p. 82
^ a b Hutterer, R.; Howell, K. (2017) [errata version of 2016 assessment]. "Crocidura fuscomurina " . IUCN Red List of Threatened Species . 2016 : e.T40633A115176835. doi :10.2305/IUCN.UK.2016-3.RLTS.T40633A22293871.en .
^ a b Raese, Joshua (2004). "Crocidura leucodon " . Animal Diversity Web . University of Michigan . Archived from the original on April 1, 2023. Retrieved March 30, 2024 .
^ a b Shenbrot, G.; Hutterer, R.; Kryštufek, B.; Yigit, N.; Mitsainas, G.; Palomo, L. (2021) [amended version of 2016 assessment]. "Crocidura leucodon " . IUCN Red List of Threatened Species . 2021 : e.T29651A197500630. doi :10.2305/IUCN.UK.2021-1.RLTS.T29651A197500630.en .
^ Kingdon , p. 114
^ a b Hutterer, R. (2016). "Crocidura nigricans " . IUCN Red List of Threatened Species . 2016 : e.T41345A22310112. doi :10.2305/IUCN.UK.2016-2.RLTS.T41345A22310112.en .
^ a b Cassola, F. (2016). "Crocidura foetida " . IUCN Red List of Threatened Species . 2016 : e.T136578A22310170. doi :10.2305/IUCN.UK.2016-2.RLTS.T136578A22310170.en .
^ Kingdon , p. 61
^ a b Gerrie, R.; Kennerley, R. (2017) [errata version of 2016 assessment]. "Crocidura bottegi " . IUCN Red List of Threatened Species . 2016 : e.T40621A115175776. doi :10.2305/IUCN.UK.2016-3.RLTS.T40621A22295879.en .
^ Kingdon , p. 63
^ Kingdon , p. 96
^ a b Cassola, F. (2017) [errata version of 2016 assessment]. "Crocidura littoralis " . IUCN Red List of Threatened Species . 2016 : e.T41330A115178517. doi :10.2305/IUCN.UK.2016-3.RLTS.T41330A22305865.en .
^ a b Kennerley, R. (2018). "Crocidura picea " . IUCN Red List of Threatened Species . 2018 : e.T5636A22301535. doi :10.2305/IUCN.UK.2018-1.RLTS.T5636A22301535.en .
^ a b c Kennerley, R. (2019). "Crocidura canariensis " . IUCN Red List of Threatened Species . 2019 : e.T5560A3031266. doi :10.2305/IUCN.UK.2019-1.RLTS.T5560A3031266.en .
^ a b Kennerley, R. (2017) [errata version of 2016 assessment]. "Crocidura caspica " . IUCN Red List of Threatened Species . 2016 : e.T136444A115207731. doi :10.2305/IUCN.UK.2016-3.RLTS.T136444A22297843.en .
^ a b Johnston, C.; Smith, A. T. (2016). "Crocidura rapax " . IUCN Red List of Threatened Species . 2016 : e.T136783A22294279. doi :10.2305/IUCN.UK.2016-1.RLTS.T136783A22294279.en .
^ a b Woinarski, J.; Burbidge, A. A.; Lumsden, L. (2016). "Crocidura trichura " . IUCN Red List of Threatened Species . 2016 : e.T136379A22304640. doi :10.2305/IUCN.UK.2016-3.RLTS.T136379A22304640.en .
^ Kingdon , p. 65
^ a b Hutterer, R.; Granjon, L. (2017) [errata version of 2016 assessment]. "Crocidura cinderella " . IUCN Red List of Threatened Species . 2016 : e.T41317A115177629. doi :10.2305/IUCN.UK.2016-3.RLTS.T41317A22307375.en .
^ a b Hutterer, R. (2016). "Crocidura congobelgica " . IUCN Red List of Threatened Species . 2016 : e.T5561A22305663. doi :10.2305/IUCN.UK.2016-2.RLTS.T5561A22305663.en .
^ a b Jenkins, P.; Dando, T.; Kennerley, R. (2017). "Crocidura cranbrooki " . IUCN Red List of Threatened Species . 2017 : e.T48296479A48296547. doi :10.2305/IUCN.UK.2017-3.RLTS.T48296479A48296547.en .
^ a b Mitsainas, G.; Vohralík, V.; Kennerley, R. (2019). "Crocidura zimmermanni " . IUCN Red List of Threatened Species . 2019 : e.T5588A90818329. doi :10.2305/IUCN.UK.2019-1.RLTS.T5588A90818329.en .
^ Kingdon , p. 68
^ a b Hutterer, R.; Jenkins, P. (2017) [errata version of 2016 assessment]. "Crocidura crossei " . IUCN Red List of Threatened Species . 2016 : e.T40624A115175904. doi :10.2305/IUCN.UK.2016-3.RLTS.T40624A22296115.en .
^ Kingdon , p. 56
^ a b Hutterer, R. (2022). "Crocidura aleksandrisi " . IUCN Red List of Threatened Species . 2022 : e.T40617A22295522. doi :10.2305/IUCN.UK.2022-2.RLTS.T40617A22295522.en .
^ Kingdon , p. 70
^ a b Hutterer, R. (2016). "Crocidura denti " . IUCN Red List of Threatened Species . 2016 : e.T40626A22294411. doi :10.2305/IUCN.UK.2016-2.RLTS.T40626A22294411.en .
^ Kingdon , p. 133
^ a b Hutterer, R.; Howell, K.; Baxter, R. (2017) [errata version of 2016 assessment]. "Crocidura smithii " . IUCN Red List of Threatened Species . 2016 : e.T41358A115181556. doi :10.2305/IUCN.UK.2016-3.RLTS.T41358A22311310.en .
^ Kingdon , p. 71
^ a b Dando, T.; Kennerley, R. (2019). "Crocidura desperata " . IUCN Red List of Threatened Species . 2019 : e.T5624A22300601. doi :10.2305/IUCN.UK.2019-3.RLTS.T5624A22300601.en .
^ a b Gerrie, R.; Kennerley, R. (2016). "Crocidura dhofarensis " . IUCN Red List of Threatened Species . 2016 : e.T5599A22303111. doi :10.2305/IUCN.UK.2016-3.RLTS.T5599A22303111.en .
^ a b Gerrie, R.; Kennerley, R. (2016). "Crocidura douceti " . IUCN Red List of Threatened Species . 2016 : e.T40629A22295053. doi :10.2305/IUCN.UK.2016-3.RLTS.T40629A22295053.en .
^ a b Cassola, F. (2017) [errata version of 2016 assessment]. "Crocidura dsinezumi " . IUCN Red List of Threatened Species . 2016 : e.T40627A115176222. doi :10.2305/IUCN.UK.2016-3.RLTS.T40627A22294822.en .
^ Kingdon , p. 57
^ a b Dando, T. (2021). "Crocidura allex " . IUCN Red List of Threatened Species . 2021 : e.T5620A22301002. doi :10.2305/IUCN.UK.2021-2.RLTS.T5620A22301002.en .
^ Kingdon , p. 128
^ a b Kryštufek, B.; Kennerley, R. (2019). "Crocidura religiosa " . IUCN Red List of Threatened Species . 2019 : e.T5616A114077410. doi :10.2305/IUCN.UK.2019-1.RLTS.T5616A114077410.en .
^ a b Kennerley, R. (2017) [errata version of 2016 assessment]. "Crocidura eisentrauti " . IUCN Red List of Threatened Species . 2016 : e.T5565A115074367. doi :10.2305/IUCN.UK.2016-3.RLTS.T5565A22305483.en .
^ a b Cassola, F. (2017) [errata version of 2016 assessment]. "Crocidura elgonius " . IUCN Red List of Threatened Species . 2016 : e.T5625A115078232. doi :10.2305/IUCN.UK.2016-3.RLTS.T5625A22299870.en .
^ a b Cassola, F. (2016). "Crocidura elongata " . IUCN Red List of Threatened Species . 2016 : e.T40630A22294921. doi :10.2305/IUCN.UK.2016-2.RLTS.T40630A22294921.en .
^ a b Ceríaco, L.; Dando, T.; Kennerley, R. (2019). "Crocidura fingui " . IUCN Red List of Threatened Species . 2019 : e.T111739377A111739380. doi :10.2305/IUCN.UK.2019-1.RLTS.T111739377A111739380.en .
^ Kingdon , p. 76
^ a b Kennerley, R.; Gerrie, R. (2017) [errata version of 2016 assessment]. "Crocidura planiceps " . IUCN Red List of Threatened Species . 2016 : e.T41351A115180837. doi :10.2305/IUCN.UK.2016-3.RLTS.T41351A22310439.en .
^ Kingdon , p. 78
^ Kingdon , p. 79
^ a b Cassola, F. (2017) [errata version of 2016 assessment]. "Crocidura foxi " . IUCN Red List of Threatened Species . 2016 : e.T5629A115078628. doi :10.2305/IUCN.UK.2016-3.RLTS.T5629A22299485.en .
^ a b c Kingdon , p. 126
^ a b Cassola, F. (2017) [errata version of 2016 assessment]. "Crocidura poensis " . IUCN Red List of Threatened Species . 2016 : e.T41352A115180968. doi :10.2305/IUCN.UK.2016-3.RLTS.T41352A22310266.en .
^ a b Jenkins, P.; Dando, T.; Kennerley, R. (2019). "Crocidura gathornei " . IUCN Red List of Threatened Species . 2019 : e.T111765432A111767002. doi :10.2305/IUCN.UK.2019-1.RLTS.T111765432A111767002.en .
^ Kingdon , p. 83
^ a b Lavrenchenko, L.; Kennerley, R. (2016). "Crocidura glassi " . IUCN Red List of Threatened Species . 2016 : e.T5566A22305372. doi :10.2305/IUCN.UK.2016-3.RLTS.T5566A22305372.en .
^ a b Hutterer, R. (2017). "Crocidura gmelini " . IUCN Red List of Threatened Species . 2017 : e.T41319A22307461. doi :10.2305/IUCN.UK.2017-2.RLTS.T41319A22307461.en .
^ a b Kingdon , p. 84
^ a b Decher, J.; Dando, T.; Kennerley, R.; Hutterer, R. (2019). "Crocidura goliath " . IUCN Red List of Threatened Species . 2019 : e.T112518189A113974008. doi :10.2305/IUCN.UK.2019-3.RLTS.T112518189A113974008.en .
^ Kingdon , p. 107
^ a b Cassola, F. (2017) [errata version of 2016 assessment]. "Crocidura maurisca " . IUCN Red List of Threatened Species . 2016 : e.T41335A115179220. doi :10.2305/IUCN.UK.2016-3.RLTS.T41335A22306327.en .
^ Kingdon , p. 86
^ a b Cassola, F. (2017) [errata version of 2016 assessment]. "Crocidura grassei " . IUCN Red List of Threatened Species . 2016 : e.T5567A115074501. doi :10.2305/IUCN.UK.2016-3.RLTS.T5567A22305289.en .
^ a b Kennerley, R. (2017) [errata version of 2016 assessment]. "Crocidura grandis " . IUCN Red List of Threatened Species . 2016 : e.T5601A115076764. doi :10.2305/IUCN.UK.2016-3.RLTS.T5601A22299006.en .
^ Kingdon , p. 77
^ a b Cassola, F. (2016). "Crocidura flavescens " . IUCN Red List of Threatened Species . 2016 : e.T5628A22300161. doi :10.2305/IUCN.UK.2016-2.RLTS.T5628A22300161.en .
^ a b Knoll, Amanda (2009). "Crocidura russula " . Animal Diversity Web . University of Michigan . Archived from the original on March 14, 2023. Retrieved March 30, 2024 .
^ a b Aulagnier, S.; Hutterer, R.; Kryštufek, B.; Yigit, N.; Mitsainas, G.; Palomo, L. (2021) [amended version of 2016 assessment]. "Crocidura russula " . IUCN Red List of Threatened Species . 2021 : e.T29652A197503499. doi :10.2305/IUCN.UK.2021-1.RLTS.T29652A197503499.en .
^ a b Hutterer, R. (2016). "Crocidura greenwoodi " . IUCN Red List of Threatened Species . 2016 : e.T5632A22299804. doi :10.2305/IUCN.UK.2016-2.RLTS.T5632A22299804.en .
^ a b Lavrenchenko, L. (2017) [errata version of 2016 assessment]. "Crocidura phaeura " . IUCN Red List of Threatened Species . 2016 : e.T5580A115075271. doi :10.2305/IUCN.UK.2016-3.RLTS.T5580A22304906.en .
^ a b Jenkins, P.; Kennerley, R.; Dando, T. (2019). "Crocidura guy " . IUCN Red List of Threatened Species . 2019 : e.T48296877A48296880. doi :10.2305/IUCN.UK.2019-1.RLTS.T48296877A48296880.en .
^ a b c Kryštufek, B.; Gazzard, A. (2023). "Crocidura gueldenstaedtii " . IUCN Red List of Threatened Species . 2023 : e.T29653A221737608. doi :10.2305/IUCN.UK.2023-1.RLTS.T29653A221737608.en .
^ a b Laginha Pinto Correia, D.; Clayton, E. (2016). "Crocidura wuchihensis " . IUCN Red List of Threatened Species . 2016 : e.T136382A22304269. doi :10.2305/IUCN.UK.2016-1.RLTS.T136382A22304269.en .
^ a b Lavrenchenko, L. (2017) [errata version of 2016 assessment]. "Crocidura harenna " . IUCN Red List of Threatened Species . 2016 : e.T5633A115078795. doi :10.2305/IUCN.UK.2016-3.RLTS.T5633A22301820.en .
^ Kingdon , p. 75
^ a b Gerrie, R.; Kennerley, R. (2017) [errata version of 2016 assessment]. "Crocidura erica " . IUCN Red List of Threatened Species . 2016 : e.T5626A115078377. doi :10.2305/IUCN.UK.2016-3.RLTS.T5626A22299982.en .
^ Kingdon , p. 89
^ a b Hutterer, R.; Jenkins, P. (2017) [errata version of 2016 assessment]. "Crocidura hildegardeae " . IUCN Red List of Threatened Species . 2016 : e.T41322A115177912. doi :10.2305/IUCN.UK.2016-3.RLTS.T41322A22307568.en .
^ a b Chiozza, F. (2016). "Crocidura hilliana " . IUCN Red List of Threatened Species . 2016 : e.T136226A22295171. doi :10.2305/IUCN.UK.2016-1.RLTS.T136226A22295171.en .
^ a b Molur, S. (2016). "Crocidura horsfieldii " . IUCN Red List of Threatened Species . 2016 : e.T41324A22307854. doi :10.2305/IUCN.UK.2016-1.RLTS.T41324A22307854.en .
^ a b Hutterer, R.; Jenkins, P. (2016). "Crocidura attila " . IUCN Red List of Threatened Species . 2016 : e.T5621A22300242. doi :10.2305/IUCN.UK.2016-2.RLTS.T5621A22300242.en .
^ a b Lunde, D. P.; Ruedas, L. A.; van Strien, N. J. (2020) [errata version of 2017 assessment]. "Crocidura hutanis " . IUCN Red List of Threatened Species . 2017 : e.T136781A165117820. doi :10.2305/IUCN.UK.2017-2.RLTS.T136781A165117820.en .
^ a b Cassola, F. (2016). "Crocidura indochinensis " . IUCN Red List of Threatened Species . 2016 : e.T136733A22299360. doi :10.2305/IUCN.UK.2016-2.RLTS.T136733A22299360.en .
^ a b Kennerley, R. (2017) [errata version of 2016 assessment]. "Crocidura susiana " . IUCN Red List of Threatened Species . 2016 : e.T5618A115077828. doi :10.2305/IUCN.UK.2016-3.RLTS.T5618A22300814.en .
^ a b Hutterer, R.; Amori, G.; Kryštufek, B. (2019). "Crocidura arispa " . IUCN Red List of Threatened Species . 2019 : e.T136270A22296424. doi :10.2305/IUCN.UK.2019-1.RLTS.T136270A22296424.en .
^ Kingdon , p. 91
^ a b Oguge, N.; Hutterer, R.; Howell, K. (2017) [errata version of 2016 assessment]. "Crocidura jacksoni " . IUCN Red List of Threatened Species . 2016 : e.T41325A115178261. doi :10.2305/IUCN.UK.2016-3.RLTS.T41325A22308229.en .
^ a b Demos, T.; Kennerley, R. (2018). "Crocidura umbra " . IUCN Red List of Threatened Species . 2018 : e.T112465479A112465482. doi :10.2305/IUCN.UK.2018-1.RLTS.T112465479A112465482.en .
^ a b Chiozza, F. (2017) [errata version of 2016 assessment]. "Crocidura maxi " . IUCN Red List of Threatened Species . 2016 : e.T41336A115179357. doi :10.2305/IUCN.UK.2016-3.RLTS.T41336A22306419.en .
^ a b Molur, S. (2017) [errata version of 2016 assessment]. "Crocidura jenkinsi " . IUCN Red List of Threatened Species . 2016 : e.T5604A115077051. doi :10.2305/IUCN.UK.2016-3.RLTS.T5604A22299101.en .
^ Kingdon , p. 92
^ a b Hutterer, R.; Jenkins, P. (2017) [errata version of 2016 assessment]. "Crocidura jouvenetae " . IUCN Red List of Threatened Species . 2016 : e.T136327A115205925. doi :10.2305/IUCN.UK.2016-3.RLTS.T136327A22309239.en .
^ a b Molur, S. (2016). "Crocidura pullata " . IUCN Red List of Threatened Species . 2016 : e.T41353A22311057. doi :10.2305/IUCN.UK.2016-1.RLTS.T41353A22311057.en .
^ a b Gerrie, R.; Kennerley, R. (2017). "Crocidura katinka " . IUCN Red List of Threatened Species . 2017 : e.T136634A22303188. doi :10.2305/IUCN.UK.2017-2.RLTS.T136634A22303188.en .
^ a b Dando, T.; Kennerley, R. (2018) [errata version of 2017 assessment]. "Crocidura monax " . IUCN Red List of Threatened Species . 2017 : e.T111767309A123795933. doi :10.2305/IUCN.UK.2017-3.RLTS.T111767309A22304558.en .
^ a b Clayton, E.; Kennerley, R. (2016). "Crocidura baluensis " . IUCN Red List of Threatened Species . 2016 : e.T136726A22300422. doi :10.2305/IUCN.UK.2016-3.RLTS.T136726A22300422.en .
^ a b Kennerley, R. (2017) [errata version of 2016 assessment]. "Crocidura lanosa " . IUCN Red List of Threatened Species . 2016 : e.T5569A115074754. doi :10.2305/IUCN.UK.2016-3.RLTS.T5569A22303856.en .
^ a b Kingdon , p. 93
^ a b Kennerley, R. (2017) [errata version of 2016 assessment]. "Crocidura kivuana " . IUCN Red List of Threatened Species . 2016 : e.T5568A115074630. doi :10.2305/IUCN.UK.2016-3.RLTS.T5568A22305202.en .
^ a b Hutterer, R. (2016). "Crocidura lamottei " . IUCN Red List of Threatened Species . 2016 : e.T41326A22308096. doi :10.2305/IUCN.UK.2016-2.RLTS.T41326A22308096.en .
^ Kingdon , p. 85
^ a b Hutterer, R. (2008). "Crocidura grandiceps " . IUCN Red List of Threatened Species . 2008 : e.T41321A10443138. doi :10.2305/IUCN.UK.2008.RLTS.T41321A10443138.en .
^ a b Hutterer, R. (2016). "Crocidura latona " . IUCN Red List of Threatened Species . 2016 : e.T5570A22303960. doi :10.2305/IUCN.UK.2016-2.RLTS.T5570A22303960.en .
^ a b Cassola, F. (2016). "Crocidura watasei " . IUCN Red List of Threatened Species . 2016 : e.T136341A22310365. doi :10.2305/IUCN.UK.2016-2.RLTS.T136341A22310365.en .
^ Kingdon , p. 132
^ a b Cassola, F. (2017) [errata version of 2016 assessment]. "Crocidura silacea " . IUCN Red List of Threatened Species . 2016 : e.T41357A115181391. doi :10.2305/IUCN.UK.2016-3.RLTS.T41357A22311376.en .
^ Kingdon , p. 90
^ a b Cassola, F. (2017) [errata version of 2016 assessment]. "Crocidura hirta " . IUCN Red List of Threatened Species . 2016 : e.T41323A115178068. doi :10.2305/IUCN.UK.2016-3.RLTS.T41323A22307953.en .
^ a b Hutterer, R. (2016). "Crocidura serezkyensis " . IUCN Red List of Threatened Species . 2016 : e.T29654A22296786. doi :10.2305/IUCN.UK.2016-2.RLTS.T29654A22296786.en .
^ a b Kryštufek, B.; Gazzard, A. (2023). "Crocidura suaveolens " . IUCN Red List of Threatened Species . 2023 : e.T221735459A221735673. doi :10.2305/IUCN.UK.2023-1.RLTS.T221735459A221735673.en .
^ Kingdon , p. 72
^ a b Hutterer, R.; Howell, K.; Jenkins, P. (2017) [errata version of 2016 assessment]. "Crocidura dolichura " . IUCN Red List of Threatened Species . 2016 : e.T40628A115176367. doi :10.2305/IUCN.UK.2016-3.RLTS.T40628A22294674.en .
^ Kingdon , p. 98
^ a b Kennerley, R.; Lavrenchenko, L. (2017) [errata version of 2016 assessment]. "Crocidura lucina " . IUCN Red List of Threatened Species . 2016 : e.T5572A115075017. doi :10.2305/IUCN.UK.2016-3.RLTS.T5572A22304095.en .
^ Kingdon , p. 99
^ a b Cassola, F. (2017) [errata version of 2016 assessment]. "Crocidura ludia " . IUCN Red List of Threatened Species . 2016 : e.T5589A115075820. doi :10.2305/IUCN.UK.2016-3.RLTS.T5589A22302289.en .
^ a b Heaney, L.; Tabaranza, B. (2016). "Crocidura grayi " . IUCN Red List of Threatened Species . 2016 : e.T5602A22298900. doi :10.2305/IUCN.UK.2016-2.RLTS.T5602A22298900.en .
^ a b Plumptre, A. J.; Dando, T.; Kennerley, R. (2020) [amended version of 2019 assessment]. "Crocidura lwiroensis " . IUCN Red List of Threatened Species . 2020 : e.T112139630A166620647. doi :10.2305/IUCN.UK.2020-1.RLTS.T112139630A166620647.en .
^ a b Kennerley, R.; Lavrenchenko, L. (2017) [errata version of 2016 assessment]. "Crocidura macmillani " . IUCN Red List of Threatened Species . 2016 : e.T5634A115078974. doi :10.2305/IUCN.UK.2016-3.RLTS.T5634A22301723.en .
^ a b Kingdon , p. 104
^ a b Cassola, F. (2017) [errata version of 2016 assessment]. "Crocidura maquassiensis " . IUCN Red List of Threatened Species . 2016 : e.T5576A115075148. doi :10.2305/IUCN.UK.2016-3.RLTS.T5576A22303779.en .
^ a b Cassola, F. (2016). "Crocidura malayana " . IUCN Red List of Threatened Species . 2016 : e.T5605A22298569. doi :10.2305/IUCN.UK.2016-2.RLTS.T5605A22298569.en .
^ a b Kennerley, R. (2016). "Crocidura virgata " . IUCN Red List of Threatened Species . 2016 : e.T136602A22309454. doi :10.2305/IUCN.UK.2016-3.RLTS.T136602A22309454.en .
^ a b Demos, T.; Kennerley, R. (2019). "Crocidura manengubae " . IUCN Red List of Threatened Species . 2019 : e.T5575A22303668. doi :10.2305/IUCN.UK.2019-1.RLTS.T5575A22303668.en .
^ a b Kingdon , p. 100
^ a b Cassola, F. (2017) [errata version of 2016 assessment]. "Crocidura lusitania " . IUCN Red List of Threatened Species . 2016 : e.T41332A115178805. doi :10.2305/IUCN.UK.2016-3.RLTS.T41332A22306053.en .
^ a b Giarla, T.; Dando, T.; Kennerley, R. (2017). "Crocidura mdumai " . IUCN Red List of Threatened Species . 2017 : e.T112503346A112503349. doi :10.2305/IUCN.UK.2017-3.RLTS.T112503346A112503349.en .
^ a b Heaney, L. (2016). "Crocidura beatus " . IUCN Red List of Threatened Species . 2016 : e.T5597A22302943. doi :10.2305/IUCN.UK.2016-2.RLTS.T5597A22302943.en .
^ a b Gerrie, R.; Kennerley, R. (2016). "Crocidura mindorus " . IUCN Red List of Threatened Species . 2016 : e.T48269124A54823641. doi :10.2305/IUCN.UK.2016-3.RLTS.T48269124A54823641.en .
^ a b Hutterer, R. (2019). "Crocidura montis " . IUCN Red List of Threatened Species . 2019 : e.T41338A22306657. doi :10.2305/IUCN.UK.2019-1.RLTS.T41338A22306657.en .
^ a b Cassola, F. (2017) [errata version of 2016 assessment]. "Crocidura luna " . IUCN Red List of Threatened Species . 2016 : e.T41331A115178646. doi :10.2305/IUCN.UK.2016-3.RLTS.T41331A22305942.en .
^ a b Gerrie, R.; Kennerley, R. (2017) [errata version of 2016 assessment]. "Crocidura musseri " . IUCN Red List of Threatened Species . 2016 : e.T136711A115211595. doi :10.2305/IUCN.UK.2016-3.RLTS.T136711A22298834.en .
^ a b Dando, T.; Kennerley, R. (2019). "Crocidura nimbasilvanus " . IUCN Red List of Threatened Species . 2019 : e.T112519468A112519471. doi :10.2305/IUCN.UK.2019-3.RLTS.T112519468A112519471.en .
^ a b Dando, T.; Kennerley, R. (2020) [amended version of 2019 assessment]. "Crocidura munissii " . IUCN Red List of Threatened Species . 2020 : e.T112503202A166619293. doi :10.2305/IUCN.UK.2020-1.RLTS.T112503202A166619293.en .
^ a b Kingdon , p. 134
^ a b Kerbis Peterhans, J.; Dieterlen, F.; Hutterer, R. (2008). "Crocidura stenocephala " . IUCN Red List of Threatened Species . 2008 : e.T5582A11371843. doi :10.2305/IUCN.UK.2008.RLTS.T5582A11371843.en .
^ a b Hutterer, R.; Shenbrot, G. (2017). "Crocidura ramona " . IUCN Red List of Threatened Species . 2017 : e.T136722A89475013. doi :10.2305/IUCN.UK.2017-2.RLTS.T136722A89475013.en .
^ a b Clayton, E.; Kennerley, R. (2018). "Crocidura negrina " . IUCN Red List of Threatened Species . 2018 : e.T5609A22298060. doi :10.2305/IUCN.UK.2018-1.RLTS.T5609A22298060.en .
^ a b Molur, S. (2017) [errata version of 2016 assessment]. "Crocidura nicobarica " . IUCN Red List of Threatened Species . 2016 : e.T5610A115077236. doi :10.2305/IUCN.UK.2016-3.RLTS.T5610A22297919.en .
^ Kingdon , p. 113
^ a b Hutterer, R.; Jenkins, P. (2017) [errata version of 2016 assessment]. "Crocidura nigeriae " . IUCN Red List of Threatened Species . 2016 : e.T41344A115179933. doi :10.2305/IUCN.UK.2016-3.RLTS.T41344A22307167.en .
^ a b Hutterer, R. (2008). "Crocidura nimbae " . IUCN Red List of Threatened Species . 2008 : e.T5578A11354307. doi :10.2305/IUCN.UK.2008.RLTS.T5578A11354307.en .
^ Kingdon , p. 117
^ a b Gerrie, R.; Kennerley, R. (2017). "Crocidura niobe " . IUCN Red List of Threatened Species . 2017 : e.T41347A22309898. doi :10.2305/IUCN.UK.2017-2.RLTS.T41347A22309898.en .
^ a b Amori, G.; Hutterer, R.; Kryštufek, B.; Yigit, N. (2022). "Crocidura pachyura " . IUCN Red List of Threatened Species . 2022 : e.T40623A22296091. doi :10.2305/IUCN.UK.2022-2.RLTS.T40623A22296091.en .
^ Kingdon , p. 103
^ a b Jenkins, P. (2017) [errata version of 2016 assessment]. "Crocidura macowi " . IUCN Red List of Threatened Species . 2016 : e.T5635A115079098. doi :10.2305/IUCN.UK.2016-3.RLTS.T5635A22301650.en .
^ a b Kennerley, R. (2017) [errata version of 2016 assessment]. "Crocidura orientalis " . IUCN Red List of Threatened Species . 2016 : e.T5611A115077427. doi :10.2305/IUCN.UK.2016-3.RLTS.T5611A22298371.en .
^ a b Cassola, F. (2016). "Crocidura palawanensis " . IUCN Red List of Threatened Species . 2016 : e.T5613A22297590. doi :10.2305/IUCN.UK.2016-2.RLTS.T5613A22297590.en .
^ a b Kennerley, R. (2017) [errata version of 2016 assessment]. "Crocidura pergrisea " . IUCN Red List of Threatened Species . 2016 : e.T5615A115077570. doi :10.2305/IUCN.UK.2016-3.RLTS.T5615A22297761.en .
^ a b Dando, T.; Kennerley, R. (2019). "Crocidura panayensis " . IUCN Red List of Threatened Species . 2019 : e.T48297786A48297816. doi :10.2305/IUCN.UK.2019-1.RLTS.T48297786A48297816.en .
^ a b Ruedas, L. (2016). "Crocidura negligens " . IUCN Red List of Threatened Species . 2016 : e.T136786A22296214. doi :10.2305/IUCN.UK.2016-1.RLTS.T136786A22296214.en .
^ a b Engelbrektsson, P.; Kennerley, R. (2016). "Crocidura phanluongi " . IUCN Red List of Threatened Species . 2016 : e.T45954204A45973011. doi :10.2305/IUCN.UK.2016-1.RLTS.T45954204A45973011.en .
^ a b Engelbrektsson, P. (2016). "Crocidura phuquocensis " . IUCN Red List of Threatened Species . 2016 : e.T45954289A45973016. doi :10.2305/IUCN.UK.2016-1.RLTS.T45954289A45973016.en .
^ Kingdon , p. 124
^ a b Gerrie, R.; Kennerley, R. (2017) [errata version of 2016 assessment]. "Crocidura pitmani " . IUCN Red List of Threatened Species . 2016 : e.T5637A115079206. doi :10.2305/IUCN.UK.2016-3.RLTS.T5637A22301462.en .
^ a b Kennerley, R. (2017) [errata version of 2016 assessment]. "Crocidura polia " . IUCN Red List of Threatened Species . 2016 : e.T5591A115075965. doi :10.2305/IUCN.UK.2016-3.RLTS.T5591A22302474.en .
^ Kingdon , p. 127
^ a b Kennerley, R. (2017) [errata version of 2016 assessment]. "Crocidura raineyi " . IUCN Red List of Threatened Species . 2016 : e.T5592A115076066. doi :10.2305/IUCN.UK.2016-3.RLTS.T5592A22302388.en .
^ Kingdon , p. 69
^ a b c Baxter, R.; Hutterer, R.; Griffin, M.; Howell, K. (2017) [errata version of 2016 assessment]. "Crocidura cyanea " . IUCN Red List of Threatened Species . 2016 : e.T40625A115176043. doi :10.2305/IUCN.UK.2016-3.RLTS.T40625A22294530.en .
^ a b Hutterer, R.; Kerbis Peterhans, J. (2017) [errata version of 2016 assessment]. "Crocidura roosevelti " . IUCN Red List of Threatened Species . 2016 : e.T41355A115181119. doi :10.2305/IUCN.UK.2016-3.RLTS.T41355A22310848.en .
^ a b Laginha Pinto Correia, D. (2016). "Crocidura orii " . IUCN Red List of Threatened Species . 2016 : e.T5590A22302169. doi :10.2305/IUCN.UK.2016-1.RLTS.T5590A22302169.en .
^ a b Jenkins, P.; Dando, T.; Kennerley, R. (2019). "Crocidura sapaensis " . IUCN Red List of Threatened Species . 2019 : e.T112465460A112465464. doi :10.2305/IUCN.UK.2019-1.RLTS.T112465460A112465464.en .
^ a b Aulagnier, S.; Gerrie, R.; Kennerley, R. (2019). "Crocidura tarfayensis " . IUCN Red List of Threatened Species . 2019 : e.T41361A22308881. doi :10.2305/IUCN.UK.2019-1.RLTS.T41361A22308881.en .
^ Kingdon , p. 121
^ a b Cassola, F. (2017) [errata version of 2016 assessment]. "Crocidura pasha " . IUCN Red List of Threatened Species . 2016 : e.T41350A115180721. doi :10.2305/IUCN.UK.2016-3.RLTS.T41350A22310542.en .
^ Kingdon , p. 112
^ a b Cassola, F. (2017) [errata version of 2016 assessment]. "Crocidura nanilla " . IUCN Red List of Threatened Species . 2016 : e.T41342A115179768. doi :10.2305/IUCN.UK.2016-3.RLTS.T41342A97250815.en .
^ a b Cassola, F. (2017) [errata version of 2016 assessment]. "Crocidura viaria " . IUCN Red List of Threatened Species . 2016 : e.T41364A115181785. doi :10.2305/IUCN.UK.2016-3.RLTS.T41364A22308708.en .
^ a b Hutterer, R.; Jenkins, P. (2017) [errata version of 2016 assessment]. "Crocidura fulvastra " . IUCN Red List of Threatened Species . 2016 : e.T40632A115176689. doi :10.2305/IUCN.UK.2016-3.RLTS.T40632A22295294.en .
^ Kingdon , p. 97
^ a b Gerrie, R.; Kennerley, R. (2017) [errata version of 2016 assessment]. "Crocidura longipes " . IUCN Red List of Threatened Species . 2016 : e.T5571A115074897. doi :10.2305/IUCN.UK.2016-3.RLTS.T5571A22304019.en .
^ a b Stubbe, M.; Samiya, R.; Ariunbold, J.; Buuveibaatar, V.; Dorjderem, S.; Monkhzul, Ts.; Otgonbaatar, M.; Tsogbadrakh, M. (2017) [errata version of 2016 assessment]. "Crocidura sibirica " . IUCN Red List of Threatened Species . 2016 : e.T41356A115181252. doi :10.2305/IUCN.UK.2016-3.RLTS.T41356A22310743.en .
^ a b Dando, T.; Kennerley, R. (2020) [amended version of 2019 assessment]. "Crocidura ninoyi " . IUCN Red List of Threatened Species . 2020 : e.T48269338A166612644. doi :10.2305/IUCN.UK.2020-1.RLTS.T48269338A166612644.en .
^ a b Giovanni Amori. (2016). "Crocidura sicula " . IUCN Red List of Threatened Species . 2016 : e.T29655A2791535. doi :10.2305/IUCN.UK.2016-2.RLTS.T29655A2791535.en .
^ a b Meegaskumbura, S.; Meegaskumbura, M. (2008). "Crocidura hikmiya " . IUCN Red List of Threatened Species . 2008 : e.T136596A4316355. doi :10.2305/IUCN.UK.2008.RLTS.T136596A4316355.en .
^ Kingdon , p. 120
^ a b Kennerley, R. (2016). "Crocidura fumosa " . IUCN Red List of Threatened Species . 2016 : e.T5630A22299610. doi :10.2305/IUCN.UK.2016-3.RLTS.T5630A22299610.en .
^ a b Engelbrektsson, P. (2016). "Crocidura sokolovi " . IUCN Red List of Threatened Species . 2016 : e.T45954331A45973021. doi :10.2305/IUCN.UK.2016-1.RLTS.T45954331A45973021.en .
^ a b Kingdon , p. 111
^ a b Cassola, F. (2019). "Crocidura nana " . IUCN Red List of Threatened Species . 2019 : e.T41341A22306927. doi :10.2305/IUCN.UK.2019-1.RLTS.T41341A22306927.en .
^ a b Cassola, F. (2017) [errata version of 2016 assessment]. "Crocidura somalica " . IUCN Red List of Threatened Species . 2016 : e.T41359A115181668. doi :10.2305/IUCN.UK.2016-3.RLTS.T41359A22311245.en .
^ a b Molur, S. (2017) [errata version of 2016 assessment]. "Crocidura fuliginosa " . IUCN Red List of Threatened Species . 2016 : e.T40631A115176525. doi :10.2305/IUCN.UK.2016-3.RLTS.T40631A22295396.en .
^ a b Dando, T.; Kennerley, R. (2019). "Crocidura miya " . IUCN Red List of Threatened Species . 2019 : e.T5608A22298694. doi :10.2305/IUCN.UK.2019-3.RLTS.T5608A22298694.en .
^ a b Chiozza, F. (2016). "Crocidura lea " . IUCN Red List of Threatened Species . 2016 : e.T41328A22308315. doi :10.2305/IUCN.UK.2016-2.RLTS.T41328A22308315.en .
^ a b Chiozza, F. (2016). "Crocidura levicula " . IUCN Red List of Threatened Species . 2016 : e.T41329A22305755. doi :10.2305/IUCN.UK.2016-2.RLTS.T41329A22305755.en .
^ a b Cassola, F. (2016). "Crocidura rhoditis " . IUCN Red List of Threatened Species . 2016 : e.T41354A22310945. doi :10.2305/IUCN.UK.2016-2.RLTS.T41354A22310945.en .
^ a b Cassola, F. (2016). "Crocidura lepidura " . IUCN Red List of Threatened Species . 2016 : e.T136792A22296335. doi :10.2305/IUCN.UK.2016-2.RLTS.T136792A22296335.en .
^ a b Cassola, F. (2016). "Crocidura paradoxura " . IUCN Red List of Threatened Species . 2016 : e.T5614A22297493. doi :10.2305/IUCN.UK.2016-2.RLTS.T5614A22297493.en .
^ a b Chiozza, F. (2016). "Crocidura monticola " . IUCN Red List of Threatened Species . 2016 : e.T41337A22306541. doi :10.2305/IUCN.UK.2016-2.RLTS.T41337A22306541.en .
^ a b Cassola, F. (2017) [errata version of 2016 assessment]. "Crocidura mariquensis " . IUCN Red List of Threatened Species . 2016 : e.T41334A115179070. doi :10.2305/IUCN.UK.2016-3.RLTS.T41334A22306233.en .
^ a b Kennerley, R. (2017) [errata version of 2016 assessment]. "Crocidura thomensis " . IUCN Red List of Threatened Species . 2016 : e.T5594A115076316. doi :10.2305/IUCN.UK.2016-3.RLTS.T5594A22303259.en .
^ a b Clayton, E. (2016). "Crocidura tanakae " . IUCN Red List of Threatened Species . 2016 : e.T136238A22294077. doi :10.2305/IUCN.UK.2016-2.RLTS.T136238A22294077.en .
^ Kingdon , p. 135
^ a b Dando, T.; Kennerley, R. (2019). "Crocidura tansaniana " . IUCN Red List of Threatened Species . 2019 : e.T112513345A112705120. doi :10.2305/IUCN.UK.2019-2.RLTS.T112513345A112705120.en .
^ a b Kennerley, R. (2016). "Crocidura tarella " . IUCN Red List of Threatened Species . 2016 : e.T41360A22311139. doi :10.2305/IUCN.UK.2016-3.RLTS.T41360A22311139.en .
^ a b Kingdon , p. 138
^ a b Kennerley, R. (2017) [errata version of 2016 assessment]. "Crocidura telfordi " . IUCN Red List of Threatened Species . 2016 : e.T5584A115075552. doi :10.2305/IUCN.UK.2016-3.RLTS.T5584A22304467.en .
^ a b Lavrenchenko, L. (2017) [errata version of 2016 assessment]. "Crocidura thalia " . IUCN Red List of Threatened Species . 2016 : e.T5585A115075702. doi :10.2305/IUCN.UK.2016-3.RLTS.T5585A22302660.en .
^ Kingdon , p. 139
^ a b Hutterer, R. (2016). "Crocidura theresae " . IUCN Red List of Threatened Species . 2016 : e.T41362A22308991. doi :10.2305/IUCN.UK.2016-2.RLTS.T41362A22308991.en .
^ a b Demos, T.; Kennerley, R. (2019). "Crocidura brunnea " . IUCN Red List of Threatened Species . 2019 : e.T136742A22297393. doi :10.2305/IUCN.UK.2019-1.RLTS.T136742A22297393.en .
^ a b Gerrie, R.; Kennerley, R. (2017) [errata version of 2016 assessment]. "Crocidura tenuis " . IUCN Red List of Threatened Species . 2016 : e.T5619A115077931. doi :10.2305/IUCN.UK.2016-3.RLTS.T5619A22300896.en .
^ Kingdon , p. 140
^ a b Hutterer, R. (2016). "Crocidura turba " . IUCN Red List of Threatened Species . 2016 : e.T41363A22308481. doi :10.2305/IUCN.UK.2016-2.RLTS.T41363A22308481.en .
^ Kingdon , p. 131
^ a b Gerrie, R.; Kennerley, R. (2017) [errata version of 2016 assessment]. "Crocidura selina " . IUCN Red List of Threatened Species . 2016 : e.T5593A115076180. doi :10.2305/IUCN.UK.2016-3.RLTS.T5593A22303358.en .
^ a b Gerrie, R.; Kennerley, R. (2017) [errata version of 2016 assessment]. "Crocidura mutesae " . IUCN Red List of Threatened Species . 2016 : e.T41340A115179654. doi :10.2305/IUCN.UK.2016-3.RLTS.T41340A22306859.en .
^ Kingdon , p. 141
^ a b Gerrie, R.; Kennerley, R. (2017) [errata version of 2016 assessment]. "Crocidura ultima " . IUCN Red List of Threatened Species . 2016 : e.T5638A115079307. doi :10.2305/IUCN.UK.2016-3.RLTS.T5638A22301388.en .
^ Kingdon , p. 149
^ a b Gerrie, R.; Kennerley, R. (2017) [errata version of 2016 assessment]. "Crocidura zimmeri " . IUCN Red List of Threatened Species . 2016 : e.T5640A115079422. doi :10.2305/IUCN.UK.2016-3.RLTS.T5640A22301128.en .
^ Kingdon , p. 142
^ a b Giarla, T.; Dando, T.; Kennerley, R. (2018) [errata version of 2017 assessment]. "Crocidura usambarae " . IUCN Red List of Threatened Species . 2017 : e.T112515709A123796512. doi :10.2305/IUCN.UK.2017-3.RLTS.T112515709A112705149.en .
^ a b c Cassola, F. (2017) [errata version of 2016 assessment]. "Crocidura lasiura " . IUCN Red List of Threatened Species . 2016 : e.T41327A115178387. doi :10.2305/IUCN.UK.2016-3.RLTS.T41327A22308395.en .
^ a b Cassola, F. (2017) [errata version of 2016 assessment]. "Crocidura voi " . IUCN Red List of Threatened Species . 2016 : e.T41365A115182001. doi :10.2305/IUCN.UK.2016-3.RLTS.T41365A22309320.en .
^ a b Chiozza, F. (2016). "Crocidura vorax " . IUCN Red List of Threatened Species . 2016 : e.T136454A22301948. doi :10.2305/IUCN.UK.2016-2.RLTS.T136454A22301948.en .
^ Kingdon , p. 110
^ a b Hutterer, R.; Howell, K. (2017) [errata version of 2016 assessment]. "Crocidura muricauda " . IUCN Red List of Threatened Species . 2016 : e.T41339A115179514. doi :10.2305/IUCN.UK.2016-3.RLTS.T41339A22306757.en .
^ Kingdon , p. 118
^ a b Nicolas, V.; Dando, T.; Kennerley, R. (2018) [errata version of 2017 assessment]. "Crocidura obscurior " . IUCN Red List of Threatened Species . 2017 : e.T111754705A123795806. doi :10.2305/IUCN.UK.2017-3.RLTS.T111754705A111754743.en .
^ Kingdon , p. 145
^ Kingdon , p. 146
^ a b Kingdon , p. 147
^ a b Hutterer, R. (2016). "Crocidura xantippe " . IUCN Red List of Threatened Species . 2016 : e.T5639A22301319. doi :10.2305/IUCN.UK.2016-2.RLTS.T5639A22301319.en .
^ a b Cassola, F. (2017) [errata version of 2016 assessment]. "Crocidura yankariensis " . IUCN Red List of Threatened Species . 2016 : e.T41367A115182133. doi :10.2305/IUCN.UK.2016-3.RLTS.T41367A22309073.en .
^ Kingdon , p. 148
^ a b Molur, S. (2017) [errata version of 2016 assessment]. "Crocidura zarudnyi " . IUCN Red List of Threatened Species . 2016 : e.T41369A115182396. doi :10.2305/IUCN.UK.2016-3.RLTS.T41369A22309535.en .
^ a b c Nowak , p. 227
^ a b c Tsytsulina, K.; Formozov, N.; Sheftel, B. (2017) [errata version of 2016 assessment]. "Diplomesodon pulchellum " . IUCN Red List of Threatened Species . 2016 : e.T41448A115186837. doi :10.2305/IUCN.UK.2016-3.RLTS.T41448A22293795.en .
^ a b c Nowak , p. 226
^ a b de A. Goonatilake, W. L. D. P. T. S.; Molur, S. (2020). "Feroculus feroculus " . IUCN Red List of Threatened Species . 2020 : e.T8553A22292753. doi :10.2305/IUCN.UK.2020-2.RLTS.T8553A22292753.en .
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^ a b Kennerley, R. (2016). "Suncus aequatorius " . IUCN Red List of Threatened Species . 2016 : e.T136224A22289522. doi :10.2305/IUCN.UK.2016-3.RLTS.T136224A22289522.en .
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^ a b Engelbrektsson, P. (2016). "Sylvisorex silvanorum " . IUCN Red List of Threatened Species . 2016 : e.T45954439A45973071. doi :10.2305/IUCN.UK.2016-1.RLTS.T45954439A45973071.en .
^ a b c d e f g h i j k l m n o Kingdon , p. 186
^ a b Kennerley, R. (2019). "Sylvisorex isabellae " . IUCN Red List of Threatened Species . 2019 : e.T21220A22290742. doi :10.2305/IUCN.UK.2019-3.RLTS.T21220A22290742.en .
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^ a b Kennerley, R. (2017) [errata version of 2016 assessment]. "Sylvisorex camerunensis " . IUCN Red List of Threatened Species . 2016 : e.T45051A115202124. doi :10.2305/IUCN.UK.2016-3.RLTS.T45051A22290959.en .
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^ a b Engelbrektsson, P. (2016). "Sylvisorex akaibei " . IUCN Red List of Threatened Species . 2016 : e.T45954406A45973066. doi :10.2305/IUCN.UK.2016-1.RLTS.T45954406A45973066.en .
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^ a b Hutterer, R. (2016). "Sylvisorex granti " . IUCN Red List of Threatened Species . 2016 : e.T41443A22291731. doi :10.2305/IUCN.UK.2016-2.RLTS.T41443A22291731.en .
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^ a b Cassola, F. (2017). "Sylvisorex ollula " . IUCN Red List of Threatened Species . 2017 : e.T48294467A22291264. doi :10.2305/IUCN.UK.2017-3.RLTS.T48294467A22291264.en .
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^ a b Kennerley, R. (2016). "Sylvisorex howelli " . IUCN Red List of Threatened Species . 2016 : e.T21216A22290854. doi :10.2305/IUCN.UK.2016-3.RLTS.T21216A22290854.en .
^ a b Cassola, F. (2017). "Sylvisorex johnstoni " . IUCN Red List of Threatened Species . 2017 : e.T41444A22291821. doi :10.2305/IUCN.UK.2017-2.RLTS.T41444A22291821.en .
^ a b Gerrie, R.; Kennerley, R. (2016). "Sylvisorex konganensis " . IUCN Red List of Threatened Species . 2016 : e.T45050A22291359. doi :10.2305/IUCN.UK.2016-3.RLTS.T45050A22291359.en .
^ a b Gerrie, R.; Kennerley, R. (2017) [errata version of 2016 assessment]. "Sylvisorex oriundus " . IUCN Red List of Threatened Species . 2016 : e.T21222A115161223. doi :10.2305/IUCN.UK.2016-3.RLTS.T21222A22291447.en .
^ a b Kingdon , p. 193
^ a b Kennerley, R. (2019). "Sylvisorex lunaris " . IUCN Red List of Threatened Species . 2019 : e.T41445A22291627. doi :10.2305/IUCN.UK.2019-1.RLTS.T41445A22291627.en .
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^ a b Kennerley, R. (2017) [errata version of 2016 assessment]. "Sylvisorex morio " . IUCN Red List of Threatened Species . 2016 : e.T21221A115161075. doi :10.2305/IUCN.UK.2016-3.RLTS.T21221A22291520.en .
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^ a b Gerrie, R.; Kennerley, R. (2019). "Sylvisorex vulcanorum " . IUCN Red List of Threatened Species . 2019 : e.T21219A22291074. doi :10.2305/IUCN.UK.2019-1.RLTS.T21219A22291074.en .
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