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List of myotines

A photograph of a greater mouse-eared bat hanging upside-down. It is gray and small.
Greater mouse-eared bat (Myotis myotis)

Myotinae is one of the four subfamilies of Vespertilionidae, itself one of twenty families of bats in the mammalian order Chiroptera and part of the microbat suborder. A member of this subfamily is called a myotine, or a mouse-eared bat. They are found in all continents besides Antarctica, primarily in forests and caves, though some species can also be found in shrublands, grasslands, rocky areas, or deserts. They range in size from the Taiwan broad-muzzled bat, at 3 cm (1 in) plus a 3 cm (1 in) tail, to the large myotis, at 10 cm (4 in) plus a 6 cm (2 in) tail. Like all bats, myotines are capable of true and sustained flight, and have wing lengths ranging from 2 cm (1 in) to 7 cm (3 in). They are all insectivorous and eat a variety of insects and spiders,[1] but some bats with long toes which trawl for insects on top of the water, such as the long-fingered bat Daubenton's bat, Maluku myotis, Rickett's big-footed bat, and pond bat, may sometimes supplement their diet with small fish from still waters. The fish-eating bat also regularly eats fish and crustaceans as well as insects, and is the only bat species that hunts fish in the ocean.[2] Almost no myotines have population estimates, though seven species—the Atacama myotis, eastern small-footed myotis, Findley's myotis, flat-headed myotis, frosted myotis, little brown bat, and peninsular myotis—are categorized as endangered species, and two species—the Nimba myotis and Yanbaru whiskered bat—are categorized as critically endangered.

The 121 extant species of Myotinae are divided between three genera: Eudiscopus and Submyotodon with one species each, and Myotis, or the mouse-eared bats, with the other 119. A few extinct prehistoric myotine species have been discovered, though due to ongoing research and discoveries the exact number and categorization is not fixed.[3]

Conventions

IUCN Red List categories
Conservation status
 EX Extinct (0 species)
 EW Extinct in the wild (0 species)
 CR Critically Endangered (2 species)
 EN Endangered (7 species)
 VU Vulnerable (8 species)
 NT Near threatened (11 species)
 LC Least concern (65 species)
Other categories
 DD Data deficient (28 species)
 NE Not evaluated (0 species)

Conservation status codes listed follow the International Union for Conservation of Nature (IUCN) Red List of Threatened Species. Range maps are provided wherever possible; if a range map is not available, a description of the myotine's range is provided. Ranges are based on the IUCN Red List for that species unless otherwise noted. Population figures are rounded to the nearest hundred.

Classification

Myotinae, one of the four subfamilies of the family Vespertilionidae, contains 121 extant species divided into 3 genera.

Subfamily Myotinae

  • Genus Eudiscopus (disk-footed bat): one species
  • Genus Myotis (mouse-eared bats): 119 species
  • Genus Submyotodon (Taiwan broad-muzzled bat): one species

Myotines

The following classification is based on the taxonomy described by the reference work Mammal Species of the World (2005), with augmentation by generally accepted proposals made since using molecular phylogenetic analysis, as supported by both the IUCN and the American Society of Mammalogists.[4]

Genus Eudiscopus Conisbee, 1953 – one species
Common name Scientific name and subspecies Range Size and ecology IUCN status and estimated population
Disk-footed bat


E. denticulus
(Osgood, 1932)
Southeastern Asia
Map of range
Size: 4–5 cm (2–2 in), plus 3–5 cm (1–2 in) tail
3–4 cm (1–2 in) arm/wing length[5]

Habitat: Forest[6]
 LC 


Unknown Unknown[6]

Genus Myotis Kaup, 1829 – 119 species
Common name Scientific name and subspecies Range Size and ecology IUCN status and estimated population
Alcathoe bat

Brown bat

M. alcathoe
von Helversen & Heller, 2001
Europe and Turkey
Map of range
Size: 3–5 cm (1–2 in), plus 3–4 cm (1–2 in) tail
3–4 cm (1–2 in) arm/wing length[7]

Habitat: Forest[8]
 DD 


Unknown Unknown[8]

Anjouan myotis


M. anjouanensis
Dorst, 1960
Madagascar
Map of range
Size: About 5 cm (2 in), plus about 5 cm (2 in) tail
about 4 cm (2 in) arm/wing length[7]

Habitat: Unknown[9]
 DD 


Unknown Unknown[9]

Anna Tess's bat


M. annatessae
Kruskop & Borisenko, 2013
Vietnam and Laos
Map of range
Size: 3–5 cm (1–2 in), plus 3–4 cm (1–2 in) tail
3–4 cm (1–2 in) arm/wing length[10]

Habitat: Forest[11]
 DD 


Unknown Unknown[11]

Annamit myotis


M. annamiticus
Kruskop & Tsytsulina, 2001
Vietnam
Map of range
Size: About 4 cm (2 in), plus about 3 cm (1 in) tail
about 4 cm (2 in) arm/wing length[10]

Habitat: Forest and inland wetlands[12]
 DD 


Unknown Unknown[12]

Arizona myotis


M. occultus
Hollister, 1909
Mexico and southwestern United States
Map of range
Size: 4–6 cm (2–2 in), plus 2–5 cm (1–2 in) tail
3–5 cm (1–2 in) arm/wing length[5]

Habitat: Forest, rocky areas, caves, and desert[13]
 LC 


Unknown Population steady[13]

Atacama myotis


M. atacamensis
Lataste, 1892
Chile and Peru
Map of range
Size: 4–5 cm (2–2 in), plus 3–4 cm (1–2 in) tail
3–4 cm (1–2 in) arm/wing length[14]

Habitat: Shrubland, rocky areas, and desert[15]
 EN 


Unknown Unknown[15]

Australian myotis


M. australis
Dobson, 1878
Australia
Map of range
Size: Unknown[16]

Habitat: Unknown[16]
 DD 


Unknown Unknown[16]

Barbados myotis


M. nyctor
LaVal & Schwartz, 1974
Lesser Antilles in the Caribbean Size: 4–5 cm (2–2 in), plus 3–4 cm (1–2 in) tail
3–4 cm (1–2 in) arm/wing length[14]

Habitat: Forest and caves[17]
 VU 


Unknown Unknown[17]

Bechstein's bat

Brown bat

M. bechsteinii
(Kuhl, 1817)
Europe and western Asia
Map of range
Size: 3–6 cm (1–2 in), plus 3–5 cm (1–2 in) tail
3–5 cm (1–2 in) arm/wing length[18]

Habitat: Forest, shrubland, and caves[19]
 NT 


Unknown Population declining[19]

Beijing mouse-eared bat


M. pequinius
Thomas, 1908
Eastern China
Map of range
Size: About 6 cm (2 in), plus about 4 cm (2 in) tail
4–6 cm (2–2 in) arm/wing length[18]

Habitat: Forest and caves[20]
 LC 


Unknown Unknown[20]

Black myotis

Black bat

M. nigricans
Schinz, 1821

Four subspecies
  • M. n. carteri
  • M. n. extremus
  • M. n. nigricans
  • M. n. osculatii
Mexico, Central America, and South America
Map of range
Size: 3–6 cm (1–2 in), plus 2–4 cm (1–2 in) tail
3–4 cm (1–2 in) arm/wing length[7]

Habitat: Forest, savanna, shrubland, and caves[21]
 LC 


Unknown Population steady[21]

Bocharic myotis


M. bucharensis
Kuzyakin, 1950
Central Asia
Map of range
Size: 4–6 cm (2–2 in), plus 4–6 cm (2–2 in) tail
3–5 cm (1–2 in) arm/wing length[18]

Habitat: Caves[22]
 DD 


Unknown Unknown[22]

Bornean whiskered myotis


M. borneoensis
Hill & Francis, 1984
Island of Borneo (in green)
Map of range
Size: Unknown length, plus 4–5 cm (2–2 in) tail
4–5 cm (2–2 in) arm/wing length[10]

Habitat: Forest[23]
 DD 


Unknown Unknown[23]

Brandt's bat

Brown bat

M. brandtii
(Eversmann, 1845)

Two subspecies
  • M. b. brandtii
  • M. b. gracilis
Europe and western Asia (in red)
Map of range
Size: 3–5 cm (1–2 in), plus 3–5 cm (1–2 in) tail
3–4 cm (1–2 in) arm/wing length[7]

Habitat: Forest, shrubland, grassland, inland wetlands, and caves[24]
 LC 


Unknown Population steady[24]

Burmese whiskered myotis

Drawing of bat head

M. montivagus
(Dobson, 1874)
Southern Asia (in leftmost red)
Map of range
Size: Unknown length
3–5 cm (1–2 in) arm/wing length[10]

Habitat: Forest and caves[25]
 DD 


Unknown Unknown[25]

California myotis

Brown bat

M. californicus
(Audubon & Bachman, 1842)

Four subspecies
  • M. c. californicus
  • M. c. caurinus
  • M. c. mexicanus
  • M. c. stephensi
Western North America
Map of range
Size: 3–5 cm (1–2 in), plus 3–5 cm (1–2 in) tail
3–4 cm (1–2 in) arm/wing length[5]

Habitat: Forest, caves, and desert[26]
 LC 


Unknown Unknown[26]

Cape hairy bat


M. tricolor
(Temminck, 1832)
Sub-Saharan Africa
Map of range
Size: 5–7 cm (2–3 in), plus 3–6 cm (1–2 in) tail
4–6 cm (2–2 in) arm/wing length[7]

Habitat: Forest, savanna, and shrubland[27]
 LC 


Unknown Unknown[27]

Cave myotis

Brown bat

M. velifer
(Allen, 1890)

Five subspecies
  • M. v. brevis
  • M. v. grandis
  • M. v. incautus
  • M. v. magnamolaris
  • M. v. velifer
United States and Mexico
Map of range
Size: 4–6 cm (2–2 in), plus 3–6 cm (1–2 in) tail
3–5 cm (1–2 in) arm/wing length[14]

Habitat: Forest and caves[28]
 LC 


Unknown Population steady[28]

Chilean myotis

Brown bat

M. chiloensis
(Waterhouse, 1840)
Southern South America
Map of range
Size: 4–5 cm (2–2 in), plus 2–4 cm (1–2 in) tail
3–5 cm (1–2 in) arm/wing length[14]

Habitat: Rocky areas and forest[29]
 LC 


Unknown Unknown[29]

Chinese water myotis


M. laniger
Peters, 1870
Eastern Asia
Map of range
Size: 3–5 cm (1–2 in), plus 3–4 cm (1–2 in) tail
3–4 cm (1–2 in) arm/wing length[10]

Habitat: Forest and caves[30]
 LC 


Unknown Unknown[30]

Cinnamon myotis


M. fortidens
Miller & Allen, 1928

Two subspecies
  • M. f. fortidens
  • M. f. sonoriensis
Guatemala and Mexico
Map of range
Size: 4–6 cm (2–2 in), plus 3–4 cm (1–2 in) tail
3–4 cm (1–2 in) arm/wing length[5]

Habitat: Forest and caves[31]
 LC 


Unknown Unknown[31]

Cryptic myotis

Brown bat

M. crypticus
Ruedi, Ibáñez, Salicini, Juste, & Puechmaille, 2019
Southern Europe Size: 4–5 cm (2–2 in), plus 3–5 cm (1–2 in) tail
3–5 cm (1–2 in) arm/wing length[18]

Habitat: Forest, inland wetlands, and caves[32]
 NT 


Unknown Population declining[32]

Csorba's mouse-eared bat


M. csorbai
Topál, 1997
Nepal Size: Unknown[33]

Habitat: Forest[33]
 DD 


Unknown Unknown[33]

Curacao myotis


M. nesopolus
Miller, 1900

Two subspecies
  • M. n. larensis
  • M. n. nesopolus
Northern South America
Map of range
Size: About 3 cm (1 in), plus about 4 cm (2 in) tail
2–4 cm (1–2 in) arm/wing length[14]

Habitat: Forest and shrubland[34]
 LC 


Unknown Unknown[34]

Dark-nosed small-footed myotis


M. melanorhinus
Merriam, 1890
Western North America
Map of range
Size: 3–5 cm (1–2 in), plus 2–5 cm (1–2 in) tail
3–4 cm (1–2 in) arm/wing length[5]

Habitat: Caves and forest[35]
 LC 


Unknown Population steady[35]

Daubenton's bat

Brown bat

M. daubentonii
(Kuhl, 1817)

Six subspecies
  • M. d. chasanensis
  • M. d. daubentonii
  • M. d. loukashkini
  • M. d. nathalinae
  • M. d. ussuriensis
  • M. d. volgensis
Europe and Asia
Map of range
Size: 4–6 cm (2–2 in), plus 2–5 cm (1–2 in) tail
3–5 cm (1–2 in) arm/wing length[18]

Habitat: Forest, shrubland, inland wetlands, rocky areas, caves, and neritic marine[36]
 LC 


Unknown Population steady[36]

David's myotis


M. davidii
Peters, 1869
Eastern Asia
Map of range
Size: 3–5 cm (1–2 in), plus 3–5 cm (1–2 in) tail
3–4 cm (1–2 in) arm/wing length[7]

Habitat: Caves and forest[37]
 LC 


Unknown Unknown[37]

Diminutive bat


M. diminutus
Moratelli & Wilson, 2011
Ecuador and Colombia Size: About 4 cm (2 in), plus about 3 cm (1 in) tail
about 3 cm (1 in) arm/wing length[14]

Habitat: Forest[38]
 DD 


Unknown Unknown[38]

Dinelli's myotis

Brown bat

M. dinellii
Thomas, 1902
Southern South America (in green)
Map of range
Size: 4–6 cm (2–2 in), plus 2–5 cm (1–2 in) tail
3–4 cm (1–2 in) arm/wing length[14]

Habitat: Forest[39]
 LC 


Unknown Population steady[39]

Dominican myotis


M. dominicensis
Miller, 1902
Dominica and Guadeloupe in the Caribbean
Map of range
Size: 4–5 cm (2–2 in), plus 2–4 cm (1–2 in) tail
3–4 cm (1–2 in) arm/wing length[14]

Habitat: Inland wetlands and caves[40]
 VU 


Unknown Unknown[40]

Eastern long-fingered bat

Gray bat

M. macrodactylus
(Temminck, 1840)

Three subspecies
  • M. m. continentalis
  • M. m. insularis
  • M. m. macrodactylus
Eastern Asia
Map of range
Size: 4–8 cm (2–3 in), plus 2–5 cm (1–2 in) tail
3–5 cm (1–2 in) arm/wing length[18]

Habitat: Forest, inland wetlands, and caves[41]
 LC 


Unknown Unknown[41]

Eastern small-footed myotis

Brown bat

M. leibii
Audubon & Bachman, 1842
Eastern United States and Canada
Map of range
Size: 4–5 cm (2–2 in), plus 3–5 cm (1–2 in) tail
3–4 cm (1–2 in) arm/wing length[5]

Habitat: Caves, rocky areas, and forest[42]
 EN 


Unknown Population declining[42]

Eastern water bat

Gray bat

M. petax
Hollister, 1912
Eastern Asia
Map of range
Size: 4–6 cm (2–2 in), plus 2–5 cm (1–2 in) tail
3–4 cm (1–2 in) arm/wing length[18]

Habitat: Caves, inland wetlands, and forest[43]
 LC 


Unknown Population steady[43]

Elegant myotis


M. elegans
Hall, 1962
Mexico and Central America
Map of range
Size: 3–5 cm (1–2 in), plus 3–4 cm (1–2 in) tail
3–4 cm (1–2 in) arm/wing length[14]

Habitat: Forest[44]
 LC 


Unknown Unknown[44]

Escalera's bat

Drawing of bat

M. escalerai
A. Cabrera, 1904
Southwestern Europe Size: 4–5 cm (2–2 in), plus 4–5 cm (2–2 in) tail
3–5 cm (1–2 in) arm/wing length[18]

Habitat: Forest, shrubland, and caves[45]
 NT 


Unknown Population declining[45]

Far Eastern myotis


M. bombinus
Thomas, 1906

Two subspecies
  • M. b. amurensis
  • M. b. bombinus
Eastern Asia
Map of range
Size: 4–6 cm (2–2 in), plus 3–5 cm (1–2 in) tail
3–5 cm (1–2 in) arm/wing length[18]

Habitat: Forest and caves[46]
 NT 


Unknown Population declining[46]

Felten's myotis

Brown bat

M. punicus
Felten, 1977
Northern Africa and Mediterranean islands
Map of range
Size: 6–8 cm (2–3 in), plus tail
5–7 cm (2–3 in) arm/wing length[18]

Habitat: Caves, grassland, shrubland, and forest[47]
 DD 


Unknown Unknown[47]

Fish-eating bat

Brown bat

M. vivesi
Ménégaux, 1901
Western Mexico
Map of range
Size: 8–9 cm (3–4 in), plus 5–8 cm (2–3 in) tail
5–7 cm (2–3 in) arm/wing length[14]

Habitat: Rocky areas and caves[48]
 VU 


Unknown Population declining[48]

Findley's myotis


M. findleyi
Bogan, 1978
Tres Marías Islands west of Mexico
Map of range
Size: 4–5 cm (2–2 in), plus 2–4 cm (1–2 in) tail
2–4 cm (1–2 in) arm/wing length[5]

Habitat: Unknown[49]
 EN 


Unknown Population declining[49]

Flat-headed myotis

Brown bat

M. planiceps
Baker, 1955
Central Mexico
Map of range
Size: 3–6 cm (1–2 in), plus 2–3 cm (1–1 in) tail
2–3 cm (1–1 in) arm/wing length[5]

Habitat: Forest and caves[50]
 EN 


240 Population declining[50]

Fraternal myotis

Stamp with drawing of bat head

M. frater
(Allen, 1923)
Eastern Asia (in red)
Map of range
Size: 3–5 cm (1–2 in), plus 4–6 cm (2–2 in) tail
3–5 cm (1–2 in) arm/wing length[18]

Habitat: Forest and caves[51]
 LC 


Unknown Population steady[51]

Fringed long-footed myotis


M. fimbriatus
Peters, 1870
Eastern Asia
Map of range
Size: 3–6 cm (1–2 in), plus 3–5 cm (1–2 in) tail
3–5 cm (1–2 in) arm/wing length[18]

Habitat: Caves[52]
 LC 


Unknown Unknown[52]

Fringed myotis

Brown bat

M. thysanodes
Miller, 1897

Four subspecies
  • M. t. aztecus
  • M. t. pahasapensis
  • M. t. thysanodes
  • M. t. vespertinus
Western North America
Map of range
Size: 4–6 cm (2–2 in), plus 3–5 cm (1–2 in) tail
4–5 cm (2–2 in) arm/wing length[5]

Habitat: Forest, rocky areas, caves, and desert[53]
 LC 


Unknown Population steady[53]

Frosted myotis


M. pruinosus
Yoshiyuki, 1971
Japan
Map of range
Size: 3–5 cm (1–2 in), plus 3–4 cm (1–2 in) tail
3–4 cm (1–2 in) arm/wing length[10]

Habitat: Forest and caves[54]
 EN 


Unknown Unknown[54]

Geoffroy's bat

Brown bat

M. emarginatus
(Geoffroy, 1806)

Three subspecies
  • M. e. desertorum
  • M. e. emarginatus
  • M. e. turcomanicus
Europe, northern Africa, and western Asia
Map of range
Size: 4–6 cm (2–2 in), plus 3–5 cm (1–2 in) tail
3–5 cm (1–2 in) arm/wing length[7]

Habitat: Caves, grassland, and shrubland[55]
 LC 


Unknown Population steady[55]

Gomantong myotis


M. gomantongensis
Francis & Hill, 1998
Island of Borneo in Malaysia
Map of range
Size: Unknown length, plus 3–5 cm (1–2 in) tail
4–5 cm (2–2 in) arm/wing length[10]

Habitat: Caves[56]
 LC 


Unknown Unknown[56]

Gray bat

Gray bat

M. grisescens
Howell, 1909
Eastern United States
Map of range
Size: 4–6 cm (2–2 in), plus 3–5 cm (1–2 in) tail
4–5 cm (2–2 in) arm/wing length[14]

Habitat: Forest and caves[57]
 VU 


Unknown Population increasing[57]

Greater mouse-eared bat

Brown bat

M. myotis
(Borkhausen, 1797)

Two subspecies
  • M. m. macrocephalicus
  • M. m. myotis
Europe and western Asia
Map of range
Size: 6–9 cm (2–4 in), plus 4–6 cm (2–2 in) tail
5–7 cm (2–3 in) arm/wing length[18]

Habitat: Forest, shrubland, and caves[58]
 LC 


Unknown Population steady[58]

Guatemalan myotis


M. cobanensis
Goodwin, 1955
Guatemala
Map of range
Size: About 4 cm (2 in), plus about 4 cm (2 in) tail
about 4 cm (2 in) arm/wing length[14]

Habitat: Unknown[59]
 DD 


Unknown Unknown[59]

Hairy-faced bat


M. annectans
Dobson, 1871
Southern and southeastern Asia
Map of range
Size: 5–7 cm (2–3 in), plus 4–5 cm (2–2 in) tail
4–5 cm (2–2 in) arm/wing length[10]

Habitat: Forest[60]
 LC 


Unknown Unknown[60]

Hairy-legged myotis

Brown bat

M. keaysi
Allen, 1914

Two subspecies
  • M. k. keaysi
  • M. k. pilosotibialis
Mexico, Central America, and northern and western South America
Map of range
Size: 5–7 cm (2–3 in), plus 2–5 cm (1–2 in) tail
Unknown arm/wing length[5]

Habitat: Forest[61]
 LC 


Unknown Unknown[61]

Herman's myotis


M. hermani
Thomas, 1923
Indonesia
Map of range
Size: 5–6 cm (2–2 in), plus 3–6 cm (1–2 in) tail
4–6 cm (2–2 in) arm/wing length[7]

Habitat: Forest[62]
 DD 


Unknown Unknown[62]

Himalayan whiskered bat


M. siligorensis
Horsfield, 1855

Four subspecies
  • M. s. alticraniatus
  • M. s. siligorensis
  • M. s. sowerbyi
  • M. s. thaianus
Central, eastern, and southeastern Asia
Map of range
Size: Unknown length
about 3 cm (1 in) arm/wing length[10]

Habitat: Forest and caves[63]
 LC 


Unknown Unknown[63]

Hodgson's bat

Brown bats

M. formosus
(Hodgson, 1835)
Central and eastern, and southeastern Asia Size: 5–6 cm (2–2 in), plus 3–6 cm (1–2 in) tail
4–6 cm (2–2 in) arm/wing length[7]

Habitat: Forest, shrubland, grassland, and caves[64]
 NT 


Unknown Population declining[64]

Horsfield's bat

Drawing of bat

M. horsfieldii
Temminck, 1840

Five subspecies
  • M. h. deignani
  • M. h. dryas
  • M. h. horsfieldii
  • M. h. jeannei
  • M. h. peshwa
Southern and southeastern Asia
Map of range
Size: 4–6 cm (2–2 in), plus 3–5 cm (1–2 in) tail
3–5 cm (1–2 in) arm/wing length[10]

Habitat: Forest and caves[65]
 LC 


Unknown Population steady[65]

Ikonnikov's bat


M. ikonnikovi
Ogniov, 1912
Eastern Asia
Map of range
Size: 3–6 cm (1–2 in), plus 2–5 cm (1–2 in) tail
3–4 cm (1–2 in) arm/wing length[7]

Habitat: Forest, inland wetlands, and caves[66]
 LC 


Unknown Population declining[66]

Indiana bat

Brown bat

M. sodalis
Miller, 1898
Eastern United States
Map of range
Size: 4–5 cm (2–2 in), plus 2–5 cm (1–2 in) tail
3–4 cm (1–2 in) arm/wing length[5]

Habitat: Forest and caves[67]
 NT 


Unknown Population steady[67]

Indochinese mouse-eared bat


M. indochinensis
Son, Motokawa, Estók, Thong, Dang, Oshida, Csorba, Francis, Görföl, & Endō, 2013
Vietnam and China
Map of range
Size: Unknown length
4–5 cm (2–2 in) arm/wing length[10]

Habitat: Forest[68]
 DD 


Unknown Unknown[68]

Insular myotis


M. insularum
Dobson, 1878
American Samoa Size: About 4 cm (2 in), plus about 4 cm (2 in) tail
about 4 cm (2 in) arm/wing length[10]

Habitat: Unknown[69]
 DD 


Unknown Unknown[69]

Izecksohn's myotis


M. izecksohni
Moratelli, Peracchi, Dias, & de Oliveira, 2011
Brazil and Argentina
Map of range
Size: 4–7 cm (2–3 in), plus 3–5 cm (1–2 in) tail
3–4 cm (1–2 in) arm/wing length[14]

Habitat: Forest[70]
 DD 


Unknown Unknown[70]

Kashmir cave bat


M. longipes
Dobson, 1873
Southern Asia Size: 3–5 cm (1–2 in), plus 3–5 cm (1–2 in) tail
3–4 cm (1–2 in) arm/wing length[10]

Habitat: Forest and caves[71]
 DD 


Unknown Unknown[71]

Keen's myotis

Brown bat

M. keenii
(Merriam, 1895)
Western Canada and United States
Map of range
Size: 4–6 cm (2–2 in), plus 3–5 cm (1–2 in) tail
3–4 cm (1–2 in) arm/wing length[5]

Habitat: Forest and rocky areas[72]
 LC 


Unknown Population steady[72]

Kei myotis


M. stalkeri
Thomas, 1910
Indonesia
Map of range
Size: 4–8 cm (2–3 in), plus 5–6 cm (2–2 in) tail
5–6 cm (2–2 in) arm/wing length[10]

Habitat: Forest, inland wetlands, and caves[73]
 LC 


10,000–11,000 Unknown[73]

Kock's mouse-eared bat


M. dieteri
Happold, 2005
Republic of the Congo Size: About 5 cm (2 in), plus about 4 cm (2 in) tail
about 4 cm (2 in) arm/wing length[7]

Habitat: Forest and caves[74]
 DD 


Unknown Unknown[74]

Large myotis


M. chinensis
Tomes, 1857
Eastern and southeastern Asia
Map of range
Size: 9–10 cm (4–4 in), plus 5–6 cm (2–2 in) tail
6–7 cm (2–3 in) arm/wing length[18]

Habitat: Forest, inland wetlands, and caves[75]
 LC 


Unknown Unknown[75]

Large-footed bat

Brown bat

M. adversus
Horsfield, 1824

Six subspecies
  • M. a. adversus
  • M. a. carimatae
  • M. a. orientis
  • M. a. taiwanensis
  • M. a. tanimbarensis
  • M. a. wetarensis
Southeastern Asia
Map of range
Size: 4–6 cm (2–2 in), plus 3–5 cm (1–2 in) tail
3–5 cm (1–2 in) arm/wing length[10]

Habitat: Forest, inland wetlands, and caves[76]
 LC 


Unknown Population steady[76]

Large-footed myotis

Black bats

M. macropus
(Gould, 1854)
Eastern Australia Size: 3–5 cm (1–2 in), plus 3–5 cm (1–2 in) tail
3–5 cm (1–2 in) arm/wing length[18]

Habitat: Inland wetlands and caves[77]
 LC 


Unknown Unknown[77]

LaVal's myotis


M. lavali
Moratelli, Peracchi, Dias, & de Oliveira, 2011
Central and eastern South America
Map of range
Size: 4–5 cm (2–2 in), plus 3–5 cm (1–2 in) tail
3–4 cm (1–2 in) arm/wing length[14]

Habitat: Forest, savanna, and grassland[78]
 LC 


Unknown Unknown[78]

Lesser large-footed bat


M. hasseltii
Temminck, 1840

Four subspecies
  • M. h. abboti
  • M. h. continentis
  • M. h. hasseltii
  • M. h. macellus
Southern and southeastern Asia
Map of range
Size: 4–6 cm (2–2 in), plus 3–5 cm (1–2 in) tail
3–5 cm (1–2 in) arm/wing length[18]

Habitat: Forest and caves[79]
 LC 


Unknown Unknown[79]

Lesser mouse-eared bat

Brown bat

M. blythii
Tomes, 1857

Four subspecies
  • M. b. ancilla
  • M. b. blythii
  • M. b. lesviacus
  • M. b. omari
Europe and Asia
Map of range
Size: 5–8 cm (2–3 in), plus about 6 cm (2 in) tail
5–7 cm (2–3 in) arm/wing length[18]

Habitat: Shrubland, grassland, and caves[80]
 LC 


Unknown Population declining[80]

Little brown bat

Brown bat

M. lucifugus
(Conte, 1831)

Five subspecies
  • M. l. alascensis
  • M. l. carissima
  • M. l. lucifugus
  • M. l. pernox
  • M. l. relictus
United States and Canada
Map of range
Size: 3–6 cm (1–2 in), plus 2–5 cm (1–2 in) tail
3–5 cm (1–2 in) arm/wing length[5]

Habitat: Forest and caves[81]
 EN 


Unknown Population declining[81]

Long-eared myotis

Brown bat

M. evotis
(H. Allen, 1864)

Six subspecies
  • M. e. chrysonotus
  • M. e. evotis
  • M. e. jonesorum
  • M. e. micronyx
  • M. e. milleri
  • M. e. pacificus
Western North America
Map of range
Size: 4–6 cm (2–2 in), plus 3–5 cm (1–2 in) tail
3–4 cm (1–2 in) arm/wing length[5]

Habitat: Caves, rocky areas, and forest[82]
 LC 


Unknown Population steady[82]

Long-fingered bat

Black bat

M. capaccinii
Bonaparte, 1837
Southern Europe, northern Africa, and western Asia
Map of range
Size: 4–6 cm (2–2 in), plus 3–5 cm (1–2 in) tail
3–5 cm (1–2 in) arm/wing length[10]

Habitat: Shrubland, inland wetlands, and caves[83]
 VU 


Unknown Population declining[83]

Long-legged myotis

Brown bat

M. volans
H. Allen, 1866

Four subspecies
  • M. v. amotus
  • M. v. interior
  • M. v. longicrus
  • M. v. volans
Western North America
Map of range
Size: 4–7 cm (2–3 in), plus 4–5 cm (2–2 in) tail
3–5 cm (1–2 in) arm/wing length[5]

Habitat: Forest, rocky areas, caves, and desert[84]
 LC 


Unknown Population steady[84]

Long-tailed myotis


M. longicaudatus
Ogniov, 1927

Four subspecies
  • M. l. eniseensis
  • M. l. kaguyae
  • M. l. longicaudatus
Eastern Asia (excluding red)
Map of range
Size: 4–6 cm (2–2 in), plus 3–5 cm (1–2 in) tail
3–5 cm (1–2 in) arm/wing length[18]

Habitat: Forest, inland wetlands, and caves[85]
 LC 


Unknown Population steady[85]

Long-toed myotis

Brown bat

M. secundus
Ruedi, Csorba, Lin, & Chou, 2015
Taiwan Size: 3–5 cm (1–2 in), plus 3–5 cm (1–2 in) tail
3–4 cm (1–2 in) arm/wing length[10]

Habitat: Forest[86]
 LC 


Unknown Population steady[86]

Malagasy mouse-eared bat


M. goudoti
Smith, 1834
Madagascar
Map of range
Size: Unknown length
3–4 cm (1–2 in) arm/wing length[7]

Habitat: Forest, rocky areas, and caves[87]
 LC 


Unknown Unknown[87]

Malaysian whiskered myotis


M. federatus
Thomas, 1916
Malaysia (in blue)
Map of range
Size: About 5 cm (2 in), plus about 4 cm (2 in) tail
3–5 cm (1–2 in) arm/wing length[10]

Habitat: Forest[88]
 DD 


Unknown Unknown[88]

Maluku myotis


M. moluccarum
Thomas, 1915

Three subspecies
  • M. m. moluccarum
  • M. m. richardsi
  • M. m. solomonis
Indonesia
Map of range
Size: 4–6 cm (2–2 in), plus 2–5 cm (1–2 in) tail
3–5 cm (1–2 in) arm/wing length[10]

Habitat: Forest and inland wetlands[89]
 LC 


Unknown Unknown[89]

Mandelli's mouse-eared bat


M. sicarius
Thomas, 1915
India and Nepal Size: 5–6 cm (2–2 in), plus 4–6 cm (2–2 in) tail
4–6 cm (2–2 in) arm/wing length[18]

Habitat: Forest and caves[90]
 VU 


Unknown Unknown[90]

Montane myotis


M. oxyotus
Peters, 1867

Two subspecies
  • M. o. gardneri
  • M. o. oxyotus
Northern and western South America
Map of range
Size: 4–6 cm (2–2 in), plus 4–5 cm (2–2 in) tail
3–5 cm (1–2 in) arm/wing length[14]

Habitat: Forest[91]
 LC 


Unknown Unknown[91]

Morris's bat


M. morrisi
Hill, 1971
Ethiopia and Nigeria
Map of range
Size: About 5 cm (2 in), plus about 5 cm (2 in) tail
4–5 cm (2–2 in) arm/wing length[7]

Habitat: Savanna and caves[92]
 DD 


Unknown Unknown[92]

Natterer's bat

Brown bat

M. nattereri
(Kuhl, 1817)

Two subspecies
  • M. n. nattereri
  • M. n. tschuliensis
Europe and western Asia Size: 4–5 cm (2–2 in), plus 3–5 cm (1–2 in) tail
3–5 cm (1–2 in) arm/wing length[18]

Habitat: Forest, shrubland, grassland, inland wetlands, and caves[93]
 LC 


Unknown Population increasing[93]

Nepal myotis


M. nipalensis
Dobson, 1871

Three subspecies
  • M. n. nipalensis
  • M. n. przewalskii
  • M. n. transcaspicus
Central and western Asia
Map of range
Size: About 4 cm (2 in), plus about 33–5 cm (1–2 in) tail
about 3 cm (1 in) arm/wing length[7]

Habitat: Forest, shrubland, grassland, caves, and desert[94]
 LC 


Unknown Unknown[94]

Nimba myotis


M. nimbaensis
Simmons, Flanders, Fils, Parker, Suter, Bamba, Douno, Keita, Morales, & Frick, 2021
Guinea Size: 6–8 cm (2–3 in), plus 4–5 cm (2–2 in) tail
5–6 cm (2–2 in) arm/wing length[7]

Habitat: Forest, grassland, inland wetlands, caves, and shrubland[95]
 CR 


Unknown Population declining[95]

Northern long-eared bat

Brown bat

M. septentrionalis
Trouessart, 1897
Canada and eastern United States
Map of range
Size: 4–5 cm (2–2 in), plus 3–5 cm (1–2 in) tail
3–4 cm (1–2 in) arm/wing length[5]

Habitat: Forest and caves[96]
 NT 


Unknown Population declining[96]

Orange-fingered myotis


M. rufopictus
(Waterhouse, 1845)
Indonesia
Map of range
Size: About 6 cm (2 in), plus 4–6 cm (2–2 in) tail
4–6 cm (2–2 in) arm/wing length[7]

Habitat: Forest[97]
 DD 


Unknown Unknown[97]

Pallid large-footed myotis


M. macrotarsus
Waterhouse, 1845

Two subspecies
  • M. m. macrotarsus
  • M. m. saba
Indonesia
Map of range
Size: About 6 cm (2 in), plus 4–6 cm (2–2 in) tail
4–5 cm (2–2 in) arm/wing length[10]

Habitat: Caves[98]
 LC 


Unknown Population steady[98]

Peninsular myotis

Brown bat

M. peninsularis
Miller, 1898
Southern Baja California peninsula
Map of range
Size: 4–6 cm (2–2 in), plus 3–6 cm (1–2 in) tail
3–5 cm (1–2 in) arm/wing length[14]

Habitat: Forest and caves[99]
 EN 


Unknown Population declining[99]

Peters's myotis


M. ater
(Peters, 1866)

Two subspecies
  • M. a. ater
  • M. a. nugax
Southeastern Asia
Map of range
Size: 4–6 cm (2–2 in), plus 3–5 cm (1–2 in) tail
3–5 cm (1–2 in) arm/wing length[10]

Habitat: Forest[100]
 LC 


Unknown Unknown[100]

Peyton's myotis


M. peytoni
Wroughton & Ryley, 1913
Southern India (in yellow)
Map of range
Size: 5–7 cm (2–3 in), plus 4–5 cm (2–2 in) tail
4–5 cm (2–2 in) arm/wing length[10]

Habitat: Forest[101]
 DD 


Unknown Unknown[101]

Pond bat

Brown bat

M. dasycneme
(Boie, 1825)
Europe and northern Asia
Map of range
Size: 5–7 cm (2–3 in), plus 4–6 cm (2–2 in) tail
4–5 cm (2–2 in) arm/wing length[7]

Habitat: Forest, inland wetlands, and caves[102]
 NT 


Unknown Population declining[102]

Red myotis

Drawing of brown bat

M. ruber
Geoffroy, 1806
Southeastern South America
Map of range
Size: 4–5 cm (2–2 in), plus 3–5 cm (1–2 in) tail
3–5 cm (1–2 in) arm/wing length[5]

Habitat: Forest[103]
 NT 


Unknown Unknown[103]

Reddish myotis


M. soror
Ruedi, Csorba, Lin, & Chou, 2015
Taiwan Size: About 5 cm (2 in), plus about 4 cm (2 in) tail
about 4 cm (2 in) arm/wing length[18]

Habitat: Forest[104]
 DD 


Unknown Unknown[104]

Reddish-black myotis

Brown and gray bat

M. rufoniger
(Tomes, 1858)
Eastern Asia
Map of range
Size: 5–8 cm (2–3 in), plus 4–6 cm (2–2 in) tail
4–6 cm (2–2 in) arm/wing length[7]

Habitat: Forest and caves[105]
 LC 


Unknown Unknown[105]

Rickett's big-footed bat


M. pilosus
Peters, 1869
Eastern Asia
Map of range
Size: 5–7 cm (2–3 in), plus 3–6 cm (1–2 in) tail
5–7 cm (2–3 in) arm/wing length[18]

Habitat: Forest and inland wetlands[106]
 VU 


Unknown Population declining[106]

Ridley's bat


M. ridleyi
Thomas, 1898
Southeastern Asia
Map of range
Size: 3–5 cm (1–2 in), plus 2–4 cm (1–2 in) tail
2–4 cm (1–2 in) arm/wing length[10]

Habitat: Caves, inland wetlands, and forest[107]
 NT 


Unknown Population declining[107]

Riparian myotis


M. riparius
Handley, 1960
Central America and South America
Map of range
Size: 4–5 cm (2–2 in), plus 2–5 cm (1–2 in) tail
3–4 cm (1–2 in) arm/wing length[14]

Habitat: Forest[108]
 LC 


Unknown Population steady[108]

Rufous mouse-eared bat

Brown bat

M. bocagii
Peters, 1870

Three subspecies
  • M. b. bocagii
  • M. b. cupreolus
  • M. b. dogalensis
Scattered sub-Saharan Africa and Yemen
Map of range
Size: 5–6 cm (2–2 in), plus 3–5 cm (1–2 in) tail
3–5 cm (1–2 in) arm/wing length[7]

Habitat: Forest and savanna[109]
 LC 


Unknown Unknown[109]

Schaub's myotis


M. schaubi
Kormos, 1934

Two subspecies
  • M. s. araxenus
  • M. s. schaubi
Armenia and Iran Size: 4–6 cm (2–2 in), plus 4–5 cm (2–2 in) tail
3–5 cm (1–2 in) arm/wing length[18]

Habitat: Forest, shrubland, and caves[110]
 DD 


Unknown Unknown[110]

Schwartz's myotis


M. martiniquensis
LaVal, 1973
Barbados and Martinique in the Caribbean
Map of range
Size: Unknown length
3–4 cm (1–2 in) arm/wing length[14]

Habitat: Caves[111]
 NT 


Unknown Unknown[111]

Scott's mouse-eared bat


M. scotti
Thomas, 1927
Ethiopia Size: 4–6 cm (2–2 in), plus 4–5 cm (2–2 in) tail
3–5 cm (1–2 in) arm/wing length[7]

Habitat: Forest and shrubland[112]
 VU 


Unknown Unknown[112]

Siberian bat

Brown bat

M. sibiricus
(Kastschenko, 1905)
Northeastern Asia (in green)
Map of range
Size: 3–6 cm (1–2 in), plus 3–5 cm (1–2 in) tail
3–4 cm (1–2 in) arm/wing length[7]

Habitat: Forest, savanna, rocky areas, and caves[113]
 LC 


Unknown Population declining[113]

Silver-tipped myotis

Drawing of bat head

M. albescens
Geoffroy, 1806
Central America and South America
Map of range
Size: 4–5 cm (2–2 in), plus 3–5 cm (1–2 in) tail
3–4 cm (1–2 in) arm/wing length[14]

Habitat: Forest[114]
 LC 


Unknown Population steady[114]

Singapore whiskered bat


M. oreias
(Temminck, 1840)
Singapore
Map of range
Size: Unknown[115]

Habitat: Unknown[115]
 DD 


Unknown Unknown[115]

Southeastern myotis

Brown bat

M. austroriparius
(Rhoads, 1897)
Southeastern United States
Map of range
Size: 4–6 cm (2–2 in), plus 2–5 cm (1–2 in) tail
3–4 cm (1–2 in) arm/wing length[14]

Habitat: Forest and caves[116]
 LC 


Unknown Population steady[116]

Southern myotis


M. aelleni
Baud, 1979
Argentina
Map of range
Size: 4–5 cm (2–2 in), plus 2–4 cm (1–2 in) tail
3–5 cm (1–2 in) arm/wing length[14]

Habitat: Unknown[117]
 DD 


Unknown Unknown[117]

Southwestern myotis

Brown bat

M. auriculus
Baker & Stains, 1955

Two subspecies
  • M. a. apache
  • M. a. auriculus
Southern United States and Mexico
Map of range
Size: 5–6 cm (2–2 in), plus 3–5 cm (1–2 in) tail
3–4 cm (1–2 in) arm/wing length[5]

Habitat: Forest, caves, and desert[118]
 LC 


Unknown Unknown[118]

Szechwan myotis


M. altarium
Thomas, 1911
China and Thailand
Map of range
Size: 4–6 cm (2–2 in), plus 3–5 cm (1–2 in) tail
4–5 cm (2–2 in) arm/wing length[7]

Habitat: Caves[119]
 LC 


Unknown Unknown[119]

Thick-thumbed myotis


M. rosseti
Oei, 1951
Southeastern Asia
Map of range
Size: 3–5 cm (1–2 in), plus 3–5 cm (1–2 in) tail
2–4 cm (1–2 in) arm/wing length[10]

Habitat: Forest[120]
 LC 


Unknown Unknown[120]

Velvety myotis


M. simus
Thomas, 1901
Western, central, and eastern South America
Map of range
Size: 5–6 cm (2–2 in), plus 3–4 cm (1–2 in) tail
3–4 cm (1–2 in) arm/wing length[14]

Habitat: Forest[121]
 DD 


Unknown Unknown[121]

Wall-roosting mouse-eared bat

Brown bat

M. muricola
(Gray, 1846)

Seven subspecies
  • M. m. browni
  • M. m. caliginosus
  • M. m. herrei
  • M. m. moupinensis
  • M. m. muricola
  • M. m. niasensis
  • M. m. patriciae
Southeastern Asia
Map of range
Size: 4–6 cm (2–2 in), plus 3–5 cm (1–2 in) tail
3–4 cm (1–2 in) arm/wing length[10]

Habitat: Forest and caves[122]
 LC 


Unknown Population steady[122]

Weber's myotis


M. weberi
(Jentink, 1890)
Sulawesi island in Indonesia Size: About 6 cm (2 in), plus about 4 cm (2 in) tail
4–6 cm (2–2 in) arm/wing length[7]

Habitat: Forest[123]
 DD 


Unknown Unknown[123]

Welwitsch's bat

Drawing of brown bat

M. welwitschii
(Gray, 1866)
Western, eastern, and southern Africa
Map of range
Size: 5–7 cm (2–3 in), plus 5–7 cm (2–3 in) tail
5–6 cm (2–2 in) arm/wing length[7]

Habitat: Forest, savanna, shrubland, and grassland[124]
 LC 


Unknown Unknown[124]

Western small-footed bat

Brown bat

M. ciliolabrum
(Merriam, 1842)
Western North America
Map of range
Size: 3–5 cm (1–2 in), plus 2–5 cm (1–2 in) tail
3–4 cm (1–2 in) arm/wing length[5]

Habitat: Forest, rocky areas, caves, and desert[125]
 LC 


Unknown Population steady[125]

Whiskered bat

Brown bat

M. mystacinus
(Kuhl, 1817)

Three subspecies
  • M. m. caucasicus
  • M. m. mystacinus
  • M. m. occidentalis
Europe, northern Africa, and western Asia
Map of range
Size: 3–5 cm (1–2 in), plus 3–5 cm (1–2 in) tail
3–4 cm (1–2 in) arm/wing length[7]

Habitat: Desert, caves, grassland, shrubland, and forest[126]
 LC 


Unknown Unknown[126]

Yanbaru whiskered bat


M. yanbarensis
Maeda & Matsumura, 1998
Ryukyu Islands in Japan
Map of range
Size: 3–5 cm (1–2 in), plus 3–5 cm (1–2 in) tail
3–4 cm (1–2 in) arm/wing length[10]

Habitat: Forest[127]
 CR 


Unknown Population declining[127]

Yellowish myotis

Drawing ofbat head

M. levis
Geoffroy, 1806
Southeastern South America (in red)
Map of range
Size: 3–6 cm (1–2 in), plus 4–5 cm (2–2 in) tail
3–5 cm (1–2 in) arm/wing length[14]

Habitat: Forest[128]
 LC 


Unknown Unknown[128]

Yuma myotis

Brown bat

M. yumanensis
H. Allen, 1864

Six subspecies
  • M. y. lambi
  • M. y. lutosus
  • M. y. oxalis
  • M. y. saturatus
  • M. y. sociabilis
  • M. y. yumanensis
Western North America
Map of range
Size: 4–5 cm (2–2 in), plus 2–4 cm (1–2 in) tail
3–4 cm (1–2 in) arm/wing length[14]

Habitat: Forest and caves[129]
 LC 


Unknown Population steady[129]

Genus Submyotodon Ziegler, 2003 – one species
Common name Scientific name and subspecies Range Size and ecology IUCN status and estimated population
Taiwan broad-muzzled bat


S. latirostris
(Kishida, 1932)
Taiwan Size: 3–4 cm (1–2 in), plus 3–4 cm (1–2 in) tail
3–4 cm (1–2 in) arm/wing length[5]

Habitat: Forest[130]
 LC 


Unknown Unknown[130]

References

  1. ^ Nowak, pp. 184–186
  2. ^ Levin, E.; Barnea, A.; Yovel, Y.; Yom-Tov, Y. (2006). "Have introduced fish initiated piscivory among the long-fingered bat?". Mammalian Biology. 71 (3): 139–143. Bibcode:2006MamBi..71..139L. doi:10.1016/j.mambio.2006.01.002.
  3. ^ "Fossilworks: Myotis". Paleobiology Database. University of Wisconsin–Madison. Archived from the original on September 22, 2022. Retrieved November 2, 2024.
  4. ^ Wilson, Reeder, pp. 499–518
  5. ^ a b c d e f g h i j k l m n o p q r s t Chernasky; Motis; Burgin, pp. 556–557
  6. ^ a b Soisook, P.; Csorba, G.; Bumrungsri, S.; Francis, C. M.; Bates, P.; Kingston, T. (2016). "Eudiscopus denticulus". IUCN Red List of Threatened Species. 2016: e.T8168A22028419. doi:10.2305/IUCN.UK.2016-2.RLTS.T8168A22028419.en.
  7. ^ a b c d e f g h i j k l m n o p q r s t u v w x y Chernasky; Motis; Burgin, pp. 560–561
  8. ^ a b Hutson, A. M.; Paunović, M. (2016). "Myotis alcathoe". IUCN Red List of Threatened Species. 2016: e.T136680A518740. doi:10.2305/IUCN.UK.2016-2.RLTS.T136680A518740.en.
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  12. ^ a b Kruskop, S. V. (2016). "Myotis annamiticus". IUCN Red List of Threatened Species. 2016: e.T136279A22006224. doi:10.2305/IUCN.UK.2016-2.RLTS.T136279A22006224.en.
  13. ^ a b Solari, S. (2018). "Myotis occultus". IUCN Red List of Threatened Species. 2018: e.T136650A21990499. doi:10.2305/IUCN.UK.2018-2.RLTS.T136650A21990499.en.
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  15. ^ a b Vargas-Rodríguez, R.; Peñaranda, D.; Ugarte Nuñez, J.; Rodríguez-San Pedro, A.; Ossa Gomez, G.; Gatica Castro, A. (2016). "Myotis atacamensis". IUCN Red List of Threatened Species. 2016: e.T14143A22050638. doi:10.2305/IUCN.UK.2016-1.RLTS.T14143A22050638.en.
  16. ^ a b c Reardon, T. B.; Lumsden, L. F. (2020). "Myotis australis". IUCN Red List of Threatened Species. 2020: e.T14146A22060248. doi:10.2305/IUCN.UK.2020-3.RLTS.T14146A22060248.en.
  17. ^ a b Larsen, R. (2016). "Myotis nyctor". IUCN Red List of Threatened Species. 2016: e.T76435059A76435083. doi:10.2305/IUCN.UK.2016-1.RLTS.T76435059A76435083.en.
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  20. ^ a b Feng, J.; Jiang, T. L. (2019). "Myotis pequinius". IUCN Red List of Threatened Species. 2019: e.T14190A22066613. doi:10.2305/IUCN.UK.2019-3.RLTS.T14190A22066613.en.
  21. ^ a b Solari, S. (2019). "Myotis nigricans". IUCN Red List of Threatened Species. 2019: e.T14185A22066939. doi:10.2305/IUCN.UK.2019-2.RLTS.T14185A22066939.en.
  22. ^ a b Srinivasulu, B.; Srinivasulu, C. (2019). "Myotis bucharensis". IUCN Red List of Threatened Species. 2019: e.T136219A22011494. doi:10.2305/IUCN.UK.2019-3.RLTS.T136219A22011494.en.
  23. ^ a b Görföl, T.; Csorba, G. (2017). "Myotis borneoensis". IUCN Red List of Threatened Species. 2017: e.T85568289A85568292. doi:10.2305/IUCN.UK.2017-2.RLTS.T85568289A85568292.en.
  24. ^ a b Gazaryan, S.; Kruskop, S. V.; Godlevska, L. (2021) [errata version of 2020 assessment]. "Myotis brandtii". IUCN Red List of Threatened Species. 2020: e.T85566997A195857637. doi:10.2305/IUCN.UK.2020-2.RLTS.T85566997A195857637.en.
  25. ^ a b Görföl, T.; Tu, V.; Csorba, G.; Francis, C. M.; Hutson, A. M.; Bates, P. J. J.; Bumrungsri, S.; Molur, S.; Srinivasulu, C. (2020). "Myotis montivagus". IUCN Red List of Threatened Species. 2020: e.T85567622A22065126. doi:10.2305/IUCN.UK.2020-2.RLTS.T85567622A22065126.en.
  26. ^ a b Arroyo-Cabrales, J.; Perez, S. (2017). "Myotis californicus". IUCN Red List of Threatened Species. 2017: e.T14150A22061366. doi:10.2305/IUCN.UK.2017-2.RLTS.T14150A22061366.en.
  27. ^ a b Monadjem, A.; Jacobs, D. (2017). "Myotis tricolor". IUCN Red List of Threatened Species. 2017: e.T14207A22063832. doi:10.2305/IUCN.UK.2017-2.RLTS.T14207A22063832.en.
  28. ^ a b Solari, S. (2019). "Myotis velifer". IUCN Red List of Threatened Species. 2019: e.T14208A22063586. doi:10.2305/IUCN.UK.2019-1.RLTS.T14208A22063586.en.
  29. ^ a b Barquez, R.; Diaz, M. (2016). "Myotis chiloensis". IUCN Red List of Threatened Species. 2016: e.T14151A22061103. doi:10.2305/IUCN.UK.2016-3.RLTS.T14151A22061103.en.
  30. ^ a b Feng, J.; Jiang, T. L. (2019). "Myotis laniger". IUCN Red List of Threatened Species. 2019: e.T136429A21984685. doi:10.2305/IUCN.UK.2019-3.RLTS.T136429A21984685.en.
  31. ^ a b Perez, S.; de Grammont, P. C.; Cuarón, A. D. (2017). "Myotis fortidens". IUCN Red List of Threatened Species. 2017: e.T14161A22056846. doi:10.2305/IUCN.UK.2017-2.RLTS.T14161A22056846.en.
  32. ^ a b Russo, D.; Cistrone, L. (2024) [errata version of 2023 assessment]. "Myotis crypticus". IUCN Red List of Threatened Species. 2023: e.T215154989A254355251.
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  34. ^ a b Solari, S. (2016). "Myotis nesopolus". IUCN Red List of Threatened Species. 2016: e.T14184A22065759. doi:10.2305/IUCN.UK.2016-1.RLTS.T14184A22065759.en.
  35. ^ a b Arroyo-Cabrales, J.; Álvarez-Castañeda, S. T. (2017). "Myotis melanorhinus". IUCN Red List of Threatened Species. 2017: e.T136784A22033542. doi:10.2305/IUCN.UK.2017-2.RLTS.T136784A22033542.en.
  36. ^ a b Kruskop, S. V.; Godlevska, L.; Bücs, S.; Çoraman, E.; Gazaryan, S. (2021) [errata version of 2020 assessment]. "Myotis daubentonii". IUCN Red List of Threatened Species. 2020: e.T85342710A195858793. doi:10.2305/IUCN.UK.2020-2.RLTS.T85342710A195858793.en.
  37. ^ a b Jiang, T. L.; Feng, J. (2019). "Myotis davidii". IUCN Red List of Threatened Species. 2019: e.T136250A22003049. doi:10.2305/IUCN.UK.2019-3.RLTS.T136250A22003049.en.
  38. ^ a b Solari, S. (2017). "Myotis diminutus". IUCN Red List of Threatened Species. 2017: e.T88151417A88151431. doi:10.2305/IUCN.UK.2017-2.RLTS.T88151417A88151431.en.
  39. ^ a b Barquez, R.; Diaz, M. (2016). "Myotis dinellii". IUCN Red List of Threatened Species. 2016: e.T136204A22009702. doi:10.2305/IUCN.UK.2016-3.RLTS.T136204A22009702.en.
  40. ^ a b Larsen, R. (2016). "Myotis dominicensis". IUCN Red List of Threatened Species. 2016: e.T14155A22057933. doi:10.2305/IUCN.UK.2016-1.RLTS.T14155A22057933.en.
  41. ^ a b Fukui, D.; Sun, K.; Kruskop, S. V. (2019). "Myotis macrodactylus". IUCN Red List of Threatened Species. 2019: e.T14177A22065868. doi:10.2305/IUCN.UK.2019-3.RLTS.T14177A22065868.en.
  42. ^ a b Solari, S. (2018). "Myotis leibii". IUCN Red List of Threatened Species. 2018: e.T14172A22055716. doi:10.2305/IUCN.UK.2018-2.RLTS.T14172A22055716.en.
  43. ^ a b Fukui, D.; Sano, A. (2020). "Myotis petax". IUCN Red List of Threatened Species. 2020: e.T85342726A85342734. doi:10.2305/IUCN.UK.2020-2.RLTS.T85342726A85342734.en.
  44. ^ a b Miller, B.; Rodriguez, B. (2017) [errata version of 2016 assessment]. "Myotis elegans". IUCN Red List of Threatened Species. 2016: e.T14156A115121563. doi:10.2305/IUCN.UK.2016-3.RLTS.T14156A22057814.en.
  45. ^ a b Russo, D.; Cistrone, L. (2023). "Myotis escalerai". IUCN Red List of Threatened Species. 2023: e.T85733126A211003991. doi:10.2305/IUCN.UK.2023-1.RLTS.T85733126A211003991.en.
  46. ^ a b Fukui, D.; Sano, A.; Kruskop, S. V. (2019). "Myotis bombinus". IUCN Red List of Threatened Species. 2019: e.T14149A22061650. doi:10.2305/IUCN.UK.2019-3.RLTS.T14149A22061650.en.
  47. ^ a b Juste, J.; Paunović, M. (2016). "Myotis punicus". IUCN Red List of Threatened Species. 2016: e.T44864A22073410. doi:10.2305/IUCN.UK.2016-2.RLTS.T44864A22073410.en.
  48. ^ a b Arroyo-Cabrales, J.; Ospina-Garces, S. (2016). "Myotis vivesi". IUCN Red List of Threatened Species. 2016: e.T14209A22069146. doi:10.2305/IUCN.UK.2016-1.RLTS.T14209A22069146.en.
  49. ^ a b Arroyo-Cabrales, J.; Ospina-Garces, S. (2016). "Myotis findleyi". IUCN Red List of Threatened Species. 2016: e.T14159A22058800. doi:10.2305/IUCN.UK.2016-1.RLTS.T14159A22058800.en.
  50. ^ a b Arroyo-Cabrales, J.; Ospina-Garces, S. (2016). "Myotis planiceps". IUCN Red List of Threatened Species. 2016: e.T14191A22066742. doi:10.2305/IUCN.UK.2016-1.RLTS.T14191A22066742.en.
  51. ^ a b Vincenot, C. E.; Preble, J. H.; Huang, J. C. -C.; Collazo, A. M.; Kamal, A. (2021). "Myotis frater". IUCN Red List of Threatened Species. 2021: e.T85566806A22056940. doi:10.2305/IUCN.UK.2021-2.RLTS.T85566806A22056940.en.
  52. ^ a b Jiang, T. L.; Feng, J. (2019). "Myotis fimbriatus". IUCN Red List of Threatened Species. 2019: e.T85735587A22058886. doi:10.2305/IUCN.UK.2019-3.RLTS.T85735587A22058886.en.
  53. ^ a b Arroyo-Cabrales, J.; de Grammont, P. C. (2017). "Myotis thysanodes". IUCN Red List of Threatened Species. 2017: e.T14206A22063246. doi:10.2305/IUCN.UK.2017-2.RLTS.T14206A22063246.en.
  54. ^ a b Fukui, D.; Sano, A. (2021) [errata version of 2019 assessment]. "Myotis pruinosus". IUCN Red List of Threatened Species. 2019: e.T14192A209551299. doi:10.2305/IUCN.UK.2019-3.RLTS.T14192A209551299.en.
  55. ^ a b Piraccini, R. (2016). "Myotis emarginatus". IUCN Red List of Threatened Species. 2016: e.T14129A22051191. doi:10.2305/IUCN.UK.2016-2.RLTS.T14129A22051191.en.
  56. ^ a b Waldien, D. L.; Santiago, K.; Wortham, G.; Stronsick, S. (2021). "Myotis gomantongensis". IUCN Red List of Threatened Species. 2021: e.T40035A22060096. doi:10.2305/IUCN.UK.2021-3.RLTS.T40035A22060096.en.
  57. ^ a b Solari, S. (2018). "Myotis grisescens". IUCN Red List of Threatened Species. 2018: e.T14132A22051652. doi:10.2305/IUCN.UK.2018-2.RLTS.T14132A22051652.en.
  58. ^ a b Coroiu, I.; Juste, J.; Paunović, M. (2016). "Myotis myotis". IUCN Red List of Threatened Species. 2016: e.T14133A22051759. doi:10.2305/IUCN.UK.2016-2.RLTS.T14133A22051759.en.
  59. ^ a b Cajas C., J.; Miller, B. (2016). "Myotis cobanensis". IUCN Red List of Threatened Species. 2016: e.T14154A22058031. doi:10.2305/IUCN.UK.2016-2.RLTS.T14154A22058031.en.
  60. ^ a b Görföl, T.; Csorba, G.; Bates, P. J. J.; Francis, C. M.; Molur, S.; Srinivasulu, C. (2020). "Myotis annectans". IUCN Red List of Threatened Species. 2020: e.T14142A22050272. doi:10.2305/IUCN.UK.2020-2.RLTS.T14142A22050272.en.
  61. ^ a b Barquez, R.; Diaz, M. (2016). "Myotis keaysi". IUCN Red List of Threatened Species. 2016: e.T14170A22056048. doi:10.2305/IUCN.UK.2016-3.RLTS.T14170A22056048.en.
  62. ^ a b Csorba, G.; Hutson, A. M.; Kingston, T.; Bumrungsri, S.; Francis, C. M. (2016). "Myotis hermani". IUCN Red List of Threatened Species. 2016: e.T14165A22057251. doi:10.2305/IUCN.UK.2016-2.RLTS.T14165A22057251.en.
  63. ^ a b Santiago, K.; Wortham, G.; Waldien, D. L. (2021). "Myotis siligorensis". IUCN Red List of Threatened Species. 2021: e.T14203A22064839. doi:10.2305/IUCN.UK.2021-3.RLTS.T14203A22064839.en.
  64. ^ a b Huang, J. C. -C.; Csorba, G.; Chang, H.-C.; Ho, Y.-Y. (2020). "Myotis formosus". IUCN Red List of Threatened Species. 2020: e.T85736120A95642290. doi:10.2305/IUCN.UK.2020-2.RLTS.T85736120A95642290.en.
  65. ^ a b Phelps, K.; Rosell-Ambal, R. G. B.; Tabaranza, B.; Heaney, L.; Gonzalez, J. C.; Molur, S.; Srinivasulu, C. (2019). "Myotis horsfieldii". IUCN Red List of Threatened Species. 2019: e.T14166A22057415. doi:10.2305/IUCN.UK.2019-3.RLTS.T14166A22057415.en.
  66. ^ a b Zhigalin, A.; Stubbe, M.; Ariunbold, J.; Buuveibaatar, V.; Dorjderem, S.; Monkhzul, T.; Otgonbaatar, M.; Tsogbadrakh, M. (2020). "Myotis ikonnikovi". IUCN Red List of Threatened Species. 2020: e.T14168A22057122. doi:10.2305/IUCN.UK.2020-3.RLTS.T14168A22057122.en.
  67. ^ a b Arroyo-Cabrales, J.; Ospina-Garces, S. (2016). "Myotis sodalis". IUCN Red List of Threatened Species. 2016: e.T14136A22053184. doi:10.2305/IUCN.UK.2016-1.RLTS.T14136A22053184.en.
  68. ^ a b Son, N.; Görföl, T.; Csorba, G. (2019). "Myotis indochinensis". IUCN Red List of Threatened Species. 2019: e.T85342688A85342691. doi:10.2305/IUCN.UK.2019-3.RLTS.T85342688A85342691.en.
  69. ^ a b Helgen, K.; Bonaccorso, F. J. (2020). "Myotis insularum". IUCN Red List of Threatened Species. 2020: e.T14169A22055968. doi:10.2305/IUCN.UK.2020-2.RLTS.T14169A22055968.en.
  70. ^ a b Solari, S. (2017). "Myotis izecksohni". IUCN Red List of Threatened Species. 2017: e.T88151563A88151572. doi:10.2305/IUCN.UK.2017-2.RLTS.T88151563A88151572.en.
  71. ^ a b Kruskop, S. V. (2016). "Myotis longipes". IUCN Red List of Threatened Species. 2016: e.T14175A22056206. doi:10.2305/IUCN.UK.2016-2.RLTS.T14175A22056206.en.
  72. ^ a b Arroyo-Cabrales, J.; Álvarez-Castañeda, S. T. (2017). "Myotis keenii". IUCN Red List of Threatened Species. 2017: e.T14171A22055579. doi:10.2305/IUCN.UK.2017-2.RLTS.T14171A22055579.en.
  73. ^ a b Bouillard, N. (2021). "Myotis stalkeri". IUCN Red List of Threatened Species. 2021: e.T14205A22063416. doi:10.2305/IUCN.UK.2021-3.RLTS.T14205A22063416.en.
  74. ^ a b Happold. M. (2019). "Myotis dieteri". IUCN Red List of Threatened Species. 2019: e.T136678A22038629. doi:10.2305/IUCN.UK.2019-3.RLTS.T136678A22038629.en.
  75. ^ a b Jiang, T. L.; Feng, J. (2019). "Myotis chinensis". IUCN Red List of Threatened Species. 2019: e.T14152A22060946. doi:10.2305/IUCN.UK.2019-3.RLTS.T14152A22060946.en.
  76. ^ a b Bouillard, N. (2021). "Myotis adversus". IUCN Red List of Threatened Species. 2021: e.T85735326A22049231. doi:10.2305/IUCN.UK.2021-2.RLTS.T85735326A22049231.en.
  77. ^ a b Gorecki, V.; Pennay, M. (2021). "Myotis macropus". IUCN Red List of Threatened Species. 2021: e.T136697A22039960. doi:10.2305/IUCN.UK.2021-1.RLTS.T136697A22039960.en.
  78. ^ a b Solari, S. (2017). "Myotis lavali". IUCN Red List of Threatened Species. 2017: e.T88151601A88151604. doi:10.2305/IUCN.UK.2017-2.RLTS.T88151601A88151604.en.
  79. ^ a b Bates, P. J. J.; Hutson, A. M.; Carino, A.; Kingston, T.; Lee, B. P. Y-H; Maryanto, I.; Molur, S.; Srinivasulu, C. (2020). "Myotis hasseltii". IUCN Red List of Threatened Species. 2020: e.T14164A22056644. doi:10.2305/IUCN.UK.2020-2.RLTS.T14164A22056644.en.
  80. ^ a b Juste, J.; Paunović, M. (2016). "Myotis blythii". IUCN Red List of Threatened Species. 2016: e.T14124A22053297. doi:10.2305/IUCN.UK.2016-2.RLTS.T14124A22053297.en.
  81. ^ a b Solari, S. (2021) [amended version of 2018 assessment]. "Myotis lucifugus". IUCN Red List of Threatened Species. 2021: e.T14176A208031565. doi:10.2305/IUCN.UK.2021-3.RLTS.T14176A208031565.en.
  82. ^ a b Arroyo-Cabrales, J.; Álvarez-Castañeda, S. T. (2017). "Myotis evotis". IUCN Red List of Threatened Species. 2017: e.T14157A22059133. doi:10.2305/IUCN.UK.2017-2.RLTS.T14157A22059133.en.
  83. ^ a b Paunović, M. (2016). "Myotis capaccinii". IUCN Red List of Threatened Species. 2016: e.T14126A22054131. doi:10.2305/IUCN.UK.2016-2.RLTS.T14126A22054131.en.
  84. ^ a b Solari, S. (2019). "Myotis volans". IUCN Red List of Threatened Species. 2019: e.T14210A22069325. doi:10.2305/IUCN.UK.2019-1.RLTS.T14210A22069325.en.
  85. ^ a b Vincenot, C. E.; Preble, J. H.; Collazo, A. M. (2021). "Myotis longicaudatus". IUCN Red List of Threatened Species. 2021: e.T85566977A85566980. doi:10.2305/IUCN.UK.2021-2.RLTS.T85566977A85566980.en.
  86. ^ a b Ruedi, M.; Csorba, G.; Liang-Kong, L.; Cheng-Han, C. (2017). "Myotis secundus". IUCN Red List of Threatened Species. 2017: e.T85342651A85342654. doi:10.2305/IUCN.UK.2017-2.RLTS.T85342651A85342654.en.
  87. ^ a b Monadjem, A.; Razafimanahaka, J.; Ranivo, J.; Kofoky, A.; Hutson, A. M.; Cardiff, S. G.; Andriafidison, D.; Goodman, S.; Jenkins, R. K. B.; Racey, P. A.; Ratrimomanarivo, F. H. (2017). "Myotis goudoti". IUCN Red List of Threatened Species. 2017: e.T14163A22056541. doi:10.2305/IUCN.UK.2017-2.RLTS.T14163A22056541.en.
  88. ^ a b Görföl, T.; Csorba, G. (2017). "Myotis federatus". IUCN Red List of Threatened Species. 2017: e.T85568302A85568305. doi:10.2305/IUCN.UK.2017-2.RLTS.T85568302A85568305.en.
  89. ^ a b Armstrong, K. N. (2021). "Myotis moluccarum". IUCN Red List of Threatened Species. 2021: e.T136770A22033795. doi:10.2305/IUCN.UK.2021-3.RLTS.T136770A22033795.en.
  90. ^ a b Srinivasulu, B.; Srinivasulu, C. (2019). "Myotis sicarius". IUCN Red List of Threatened Species. 2019: e.T14202A22063965. doi:10.2305/IUCN.UK.2019-3.RLTS.T14202A22063965.en.
  91. ^ a b Solari, S. (2018). "Myotis oxyotus". IUCN Red List of Threatened Species. 2018: e.T14187A22067211. doi:10.2305/IUCN.UK.2018-2.RLTS.T14187A22067211.en.
  92. ^ a b Jacobs, D.; Cotterill, F. P. D.; Taylor, P. J. (2019). "Myotis morrisi". IUCN Red List of Threatened Species. 2019: e.T14182A22065314. doi:10.2305/IUCN.UK.2019-3.RLTS.T14182A22065314.en.
  93. ^ a b Russo, D.; Cistrone, L. (2023). "Myotis nattereri". IUCN Red List of Threatened Species. 2023: e.T215492021A211005466. doi:10.2305/IUCN.UK.2023-1.RLTS.T215492021A211005466.en.
  94. ^ a b Srinivasulu, B.; Srinivasulu, C. (2019). "Myotis nipalensis". IUCN Red List of Threatened Species. 2019: e.T136495A21976309. doi:10.2305/IUCN.UK.2019-3.RLTS.T136495A21976309.en.
  95. ^ a b Bakwo Fils, E. M.; Flanders, J.; Frick, W. F.; Simmons, N. (2022). "Myotis nimbaensis". IUCN Red List of Threatened Species. 2022: e.T216617275A216617367. doi:10.2305/IUCN.UK.2022-2.RLTS.T216617275A216617367.en.
  96. ^ a b Solari, S. (2018). "Myotis septentrionalis". IUCN Red List of Threatened Species. 2018: e.T14201A22064312. doi:10.2305/IUCN.UK.2018-2.RLTS.T14201A22064312.en.
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  98. ^ a b Duya, M. R.; Alvarez, J.; Fidelino, J.; Gatan-Balbas, M.; Pedregosa, M.; Veluz, M. J.; Jakosalem, P. G.; Tanalgo, K.; Garcia, J. J. L.; Cacho, M. A. (2019). "Myotis macrotarsus". IUCN Red List of Threatened Species. 2019: e.T14178A22065997. doi:10.2305/IUCN.UK.2019-3.RLTS.T14178A22065997.en.
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