List of myotines
Species in mammal subfamily Myotinae
Greater mouse-eared bat (Myotis myotis )
Myotinae is one of the four subfamilies of Vespertilionidae , itself one of twenty families of bats in the mammalian order Chiroptera and part of the microbat suborder. A member of this subfamily is called a myotine, or a mouse-eared bat. They are found in all continents besides Antarctica, primarily in forests and caves, though some species can also be found in shrublands , grasslands , rocky areas, or deserts. They range in size from the Taiwan broad-muzzled bat , at 3 cm (1 in) plus a 3 cm (1 in) tail, to the large myotis , at 10 cm (4 in) plus a 6 cm (2 in) tail. Like all bats, myotines are capable of true and sustained flight , and have wing lengths ranging from 2 cm (1 in) to 7 cm (3 in). They are all insectivorous and eat a variety of insects and spiders,[ 1] but some bats with long toes which trawl for insects on top of the water, such as the long-fingered bat Daubenton's bat , Maluku myotis , Rickett's big-footed bat , and pond bat , may sometimes supplement their diet with small fish from still waters. The fish-eating bat also regularly eats fish and crustaceans as well as insects, and is the only bat species that hunts fish in the ocean.[ 2] Almost no myotines have population estimates, though seven species—the Atacama myotis , eastern small-footed myotis , Findley's myotis , flat-headed myotis , frosted myotis , little brown bat , and peninsular myotis —are categorized as endangered species , and two species—the Nimba myotis and Yanbaru whiskered bat —are categorized as critically endangered .
The 121 extant species of Myotinae are divided between three genera : Eudiscopus and Submyotodon with one species each, and Myotis , or the mouse-eared bats, with the other 119. A few extinct prehistoric myotine species have been discovered, though due to ongoing research and discoveries the exact number and categorization is not fixed.[ 3]
Conventions
Conservation status codes listed follow the International Union for Conservation of Nature (IUCN) Red List of Threatened Species . Range maps are provided wherever possible; if a range map is not available, a description of the myotine's range is provided. Ranges are based on the IUCN Red List for that species unless otherwise noted. Population figures are rounded to the nearest hundred.
Classification
Myotinae , one of the four subfamilies of the family Vespertilionidae , contains 121 extant species divided into 3 genera .
Subfamily Myotinae
Genus Eudiscopus (disk-footed bat): one species
Genus Myotis (mouse-eared bats): 119 species
Genus Submyotodon (Taiwan broad-muzzled bat): one species
Myotines
The following classification is based on the taxonomy described by the reference work Mammal Species of the World (2005), with augmentation by generally accepted proposals made since using molecular phylogenetic analysis , as supported by both the IUCN and the American Society of Mammalogists .[ 4]
Genus Eudiscopus – Conisbee , 1953 – one species
Common name
Scientific name and subspecies
Range
Size and ecology
IUCN status and estimated population
Disk-footed bat
E. denticulus (Osgood , 1932)
Southeastern Asia
Size : 4–5 cm (2–2 in), plus 3–5 cm (1–2 in) tail 3–4 cm (1–2 in) arm/wing length[ 5] Habitat : Forest
LC
Unknown
Genus Myotis – Kaup , 1829 – 119 species
Common name
Scientific name and subspecies
Range
Size and ecology
IUCN status and estimated population
Alcathoe bat
M. alcathoe von Helversen & Heller , 2001
Europe and Turkey
Size : 3–5 cm (1–2 in), plus 3–4 cm (1–2 in) tail 3–4 cm (1–2 in) arm/wing length[ 7] Habitat : Forest[ 8]
DD
Unknown [ 8]
Anjouan myotis
M. anjouanensis Dorst , 1960
Madagascar
Size : About 5 cm (2 in), plus about 5 cm (2 in) tail about 4 cm (2 in) arm/wing length[ 7] Habitat : Unknown[ 9]
DD
Unknown [ 9]
Anna Tess's bat
M. annatessae Kruskop & Borisenko , 2013
Vietnam and Laos
Size : 3–5 cm (1–2 in), plus 3–4 cm (1–2 in) tail 3–4 cm (1–2 in) arm/wing length[ 10] Habitat : Forest[ 11]
DD
Unknown [ 11]
Annamit myotis
M. annamiticus Kruskop & Tsytsulina , 2001
Vietnam
Size : About 4 cm (2 in), plus about 3 cm (1 in) tail about 4 cm (2 in) arm/wing length[ 10] Habitat : Forest and inland wetlands[ 12]
DD
Unknown [ 12]
Arizona myotis
M. occultus Hollister , 1909
Mexico and southwestern United States
Size : 4–6 cm (2–2 in), plus 2–5 cm (1–2 in) tail 3–5 cm (1–2 in) arm/wing length[ 5] Habitat : Forest, rocky areas, caves, and desert[ 13]
LC
Unknown [ 13]
Atacama myotis
M. atacamensis Lataste , 1892
Chile and Peru
Size : 4–5 cm (2–2 in), plus 3–4 cm (1–2 in) tail 3–4 cm (1–2 in) arm/wing length[ 14] Habitat : Shrubland, rocky areas, and desert[ 15]
EN
Unknown [ 15]
Australian myotis
M. australis Dobson , 1878
Australia
Size : Unknown[ 16] Habitat : Unknown[ 16]
DD
Unknown [ 16]
Barbados myotis
M. nyctor LaVal & Schwartz , 1974
Lesser Antilles in the Caribbean
Size : 4–5 cm (2–2 in), plus 3–4 cm (1–2 in) tail 3–4 cm (1–2 in) arm/wing length[ 14] Habitat : Forest and caves[ 17]
VU
Unknown [ 17]
Bechstein's bat
M. bechsteinii (Kuhl , 1817)
Europe and western Asia
Size : 3–6 cm (1–2 in), plus 3–5 cm (1–2 in) tail 3–5 cm (1–2 in) arm/wing length[ 18] Habitat : Forest, shrubland, and caves[ 19]
NT
Unknown [ 19]
Beijing mouse-eared bat
M. pequinius Thomas , 1908
Eastern China
Size : About 6 cm (2 in), plus about 4 cm (2 in) tail 4–6 cm (2–2 in) arm/wing length[ 18] Habitat : Forest and caves[ 20]
LC
Unknown [ 20]
Black myotis
M. nigricans Schinz , 1821
M. n. carteri
M. n. extremus
M. n. nigricans
M. n. osculatii
Mexico, Central America, and South America
Size : 3–6 cm (1–2 in), plus 2–4 cm (1–2 in) tail 3–4 cm (1–2 in) arm/wing length[ 7] Habitat : Forest, savanna, shrubland, and caves[ 21]
LC
Unknown [ 21]
Bocharic myotis
M. bucharensis Kuzyakin , 1950
Central Asia
Size : 4–6 cm (2–2 in), plus 4–6 cm (2–2 in) tail 3–5 cm (1–2 in) arm/wing length[ 18] Habitat : Caves[ 22]
DD
Unknown [ 22]
Bornean whiskered myotis
M. borneoensis Hill & Francis , 1984
Island of Borneo (in green)
Size : Unknown length, plus 4–5 cm (2–2 in) tail 4–5 cm (2–2 in) arm/wing length[ 10] Habitat : Forest[ 23]
DD
Unknown [ 23]
Brandt's bat
M. brandtii (Eversmann , 1845)
M. b. brandtii
M. b. gracilis
Europe and western Asia (in red)
Size : 3–5 cm (1–2 in), plus 3–5 cm (1–2 in) tail 3–4 cm (1–2 in) arm/wing length[ 7] Habitat : Forest, shrubland, grassland, inland wetlands, and caves[ 24]
LC
Unknown [ 24]
Burmese whiskered myotis
M. montivagus (Dobson , 1874)
Southern Asia (in leftmost red)
Size : Unknown length 3–5 cm (1–2 in) arm/wing length[ 10] Habitat : Forest and caves[ 25]
DD
Unknown [ 25]
California myotis
M. californicus (Audubon & Bachman , 1842)
M. c. californicus
M. c. caurinus
M. c. mexicanus
M. c. stephensi
Western North America
Size : 3–5 cm (1–2 in), plus 3–5 cm (1–2 in) tail 3–4 cm (1–2 in) arm/wing length[ 5] Habitat : Forest, caves, and desert[ 26]
LC
Unknown [ 26]
Cape hairy bat
M. tricolor (Temminck , 1832)
Sub-Saharan Africa
Size : 5–7 cm (2–3 in), plus 3–6 cm (1–2 in) tail 4–6 cm (2–2 in) arm/wing length[ 7] Habitat : Forest, savanna, and shrubland[ 27]
LC
Unknown [ 27]
Cave myotis
M. velifer (Allen , 1890)
M. v. brevis
M. v. grandis
M. v. incautus
M. v. magnamolaris
M. v. velifer
United States and Mexico
Size : 4–6 cm (2–2 in), plus 3–6 cm (1–2 in) tail 3–5 cm (1–2 in) arm/wing length[ 14] Habitat : Forest and caves[ 28]
LC
Unknown [ 28]
Chilean myotis
M. chiloensis (Waterhouse , 1840)
Southern South America
Size : 4–5 cm (2–2 in), plus 2–4 cm (1–2 in) tail 3–5 cm (1–2 in) arm/wing length[ 14] Habitat : Rocky areas and forest[ 29]
LC
Unknown [ 29]
Chinese water myotis
M. laniger Peters , 1870
Eastern Asia
Size : 3–5 cm (1–2 in), plus 3–4 cm (1–2 in) tail 3–4 cm (1–2 in) arm/wing length[ 10] Habitat : Forest and caves[ 30]
LC
Unknown [ 30]
Cinnamon myotis
M. fortidens Miller & Allen , 1928
M. f. fortidens
M. f. sonoriensis
Guatemala and Mexico
Size : 4–6 cm (2–2 in), plus 3–4 cm (1–2 in) tail 3–4 cm (1–2 in) arm/wing length[ 5] Habitat : Forest and caves[ 31]
LC
Unknown [ 31]
Cryptic myotis
M. crypticus Ruedi , Ibáñez , Salicini , Juste , & Puechmaille , 2019
Southern Europe
Size : 4–5 cm (2–2 in), plus 3–5 cm (1–2 in) tail 3–5 cm (1–2 in) arm/wing length[ 18] Habitat : Forest, inland wetlands, and caves[ 32]
NT
Unknown [ 32]
Csorba's mouse-eared bat
M. csorbai Topál , 1997
Nepal
Size : Unknown[ 33] Habitat : Forest[ 33]
DD
Unknown [ 33]
Curacao myotis
M. nesopolus Miller , 1900
M. n. larensis
M. n. nesopolus
Northern South America
Size : About 3 cm (1 in), plus about 4 cm (2 in) tail 2–4 cm (1–2 in) arm/wing length[ 14] Habitat : Forest and shrubland[ 34]
LC
Unknown [ 34]
Dark-nosed small-footed myotis
M. melanorhinus Merriam , 1890
Western North America
Size : 3–5 cm (1–2 in), plus 2–5 cm (1–2 in) tail 3–4 cm (1–2 in) arm/wing length[ 5] Habitat : Caves and forest
LC
Unknown
Daubenton's bat
M. daubentonii (Kuhl , 1817)
M. d. chasanensis
M. d. daubentonii
M. d. loukashkini
M. d. nathalinae
M. d. ussuriensis
M. d. volgensis
Europe and Asia
Size : 4–6 cm (2–2 in), plus 2–5 cm (1–2 in) tail 3–5 cm (1–2 in) arm/wing length[ 18] Habitat : Forest, shrubland, inland wetlands, rocky areas, caves, and neritic marine [ 36]
LC
Unknown [ 36]
David's myotis
M. davidii Peters , 1869
Eastern Asia
Size : 3–5 cm (1–2 in), plus 3–5 cm (1–2 in) tail 3–4 cm (1–2 in) arm/wing length[ 7] Habitat : Caves and forest[ 37]
LC
Unknown [ 37]
Diminutive bat
M. diminutus Moratelli & Wilson , 2011
Ecuador and Colombia
Size : About 4 cm (2 in), plus about 3 cm (1 in) tail about 3 cm (1 in) arm/wing length[ 14] Habitat : Forest[ 38]
DD
Unknown [ 38]
Dinelli's myotis
M. dinellii Thomas , 1902
Southern South America (in green)
Size : 4–6 cm (2–2 in), plus 2–5 cm (1–2 in) tail 3–4 cm (1–2 in) arm/wing length[ 14] Habitat : Forest[ 39]
LC
Unknown [ 39]
Dominican myotis
M. dominicensis Miller , 1902
Dominica and Guadeloupe in the Caribbean
Size : 4–5 cm (2–2 in), plus 2–4 cm (1–2 in) tail 3–4 cm (1–2 in) arm/wing length[ 14] Habitat : Inland wetlands and caves[ 40]
VU
Unknown [ 40]
Eastern long-fingered bat
M. macrodactylus (Temminck , 1840)
M. m. continentalis
M. m. insularis
M. m. macrodactylus
Eastern Asia
Size : 4–8 cm (2–3 in), plus 2–5 cm (1–2 in) tail 3–5 cm (1–2 in) arm/wing length[ 18] Habitat : Forest, inland wetlands, and caves[ 41]
LC
Unknown [ 41]
Eastern small-footed myotis
M. leibii Audubon & Bachman , 1842
Eastern United States and Canada
Size : 4–5 cm (2–2 in), plus 3–5 cm (1–2 in) tail 3–4 cm (1–2 in) arm/wing length[ 5] Habitat : Caves, rocky areas, and forest
EN
Unknown
Eastern water bat
M. petax Hollister , 1912
Eastern Asia
Size : 4–6 cm (2–2 in), plus 2–5 cm (1–2 in) tail 3–4 cm (1–2 in) arm/wing length[ 18] Habitat : Caves, inland wetlands, and forest[ 43]
LC
Unknown [ 43]
Elegant myotis
M. elegans Hall , 1962
Mexico and Central America
Size : 3–5 cm (1–2 in), plus 3–4 cm (1–2 in) tail 3–4 cm (1–2 in) arm/wing length[ 14] Habitat : Forest[ 44]
LC
Unknown [ 44]
Escalera's bat
M. escalerai A. Cabrera , 1904
Southwestern Europe
Size : 4–5 cm (2–2 in), plus 4–5 cm (2–2 in) tail 3–5 cm (1–2 in) arm/wing length[ 18] Habitat : Forest, shrubland, and caves[ 45]
NT
Unknown [ 45]
Far Eastern myotis
M. bombinus Thomas , 1906
M. b. amurensis
M. b. bombinus
Eastern Asia
Size : 4–6 cm (2–2 in), plus 3–5 cm (1–2 in) tail 3–5 cm (1–2 in) arm/wing length[ 18] Habitat : Forest and caves[ 46]
NT
Unknown [ 46]
Felten's myotis
M. punicus Felten , 1977
Northern Africa and Mediterranean islands
Size : 6–8 cm (2–3 in), plus tail 5–7 cm (2–3 in) arm/wing length[ 18] Habitat : Caves, grassland, shrubland, and forest[ 47]
DD
Unknown [ 47]
Fish-eating bat
M. vivesi Ménégaux , 1901
Western Mexico
Size : 8–9 cm (3–4 in), plus 5–8 cm (2–3 in) tail 5–7 cm (2–3 in) arm/wing length[ 14] Habitat : Rocky areas and caves[ 48]
VU
Unknown [ 48]
Findley's myotis
M. findleyi Bogan , 1978
Tres Marías Islands west of Mexico
Size : 4–5 cm (2–2 in), plus 2–4 cm (1–2 in) tail 2–4 cm (1–2 in) arm/wing length[ 5] Habitat : Unknown[ 49]
EN
Unknown [ 49]
Flat-headed myotis
M. planiceps Baker , 1955
Central Mexico
Size : 3–6 cm (1–2 in), plus 2–3 cm (1–1 in) tail 2–3 cm (1–1 in) arm/wing length[ 5] Habitat : Forest and caves[ 50]
EN
240 [ 50]
Fraternal myotis
M. frater (Allen , 1923)
Eastern Asia (in red)
Size : 3–5 cm (1–2 in), plus 4–6 cm (2–2 in) tail 3–5 cm (1–2 in) arm/wing length[ 18] Habitat : Forest and caves[ 51]
LC
Unknown [ 51]
Fringed long-footed myotis
M. fimbriatus Peters , 1870
Eastern Asia
Size : 3–6 cm (1–2 in), plus 3–5 cm (1–2 in) tail 3–5 cm (1–2 in) arm/wing length[ 18] Habitat : Caves
LC
Unknown
Fringed myotis
M. thysanodes Miller , 1897
M. t. aztecus
M. t. pahasapensis
M. t. thysanodes
M. t. vespertinus
Western North America
Size : 4–6 cm (2–2 in), plus 3–5 cm (1–2 in) tail 4–5 cm (2–2 in) arm/wing length[ 5] Habitat : Forest, rocky areas, caves, and desert[ 53]
LC
Unknown [ 53]
Frosted myotis
M. pruinosus Yoshiyuki , 1971
Japan
Size : 3–5 cm (1–2 in), plus 3–4 cm (1–2 in) tail 3–4 cm (1–2 in) arm/wing length[ 10] Habitat : Forest and caves[ 54]
EN
Unknown [ 54]
Geoffroy's bat
M. emarginatus (Geoffroy , 1806)
M. e. desertorum
M. e. emarginatus
M. e. turcomanicus
Europe, northern Africa, and western Asia
Size : 4–6 cm (2–2 in), plus 3–5 cm (1–2 in) tail 3–5 cm (1–2 in) arm/wing length[ 7] Habitat : Caves, grassland, and shrubland[ 55]
LC
Unknown [ 55]
Gomantong myotis
M. gomantongensis Francis & Hill , 1998
Island of Borneo in Malaysia
Size : Unknown length, plus 3–5 cm (1–2 in) tail 4–5 cm (2–2 in) arm/wing length[ 10] Habitat : Caves[ 56]
LC
Unknown [ 56]
Gray bat
M. grisescens Howell , 1909
Eastern United States
Size : 4–6 cm (2–2 in), plus 3–5 cm (1–2 in) tail 4–5 cm (2–2 in) arm/wing length[ 14] Habitat : Forest and caves[ 57]
VU
Unknown [ 57]
Greater mouse-eared bat
M. myotis (Borkhausen , 1797)
M. m. macrocephalicus
M. m. myotis
Europe and western Asia
Size : 6–9 cm (2–4 in), plus 4–6 cm (2–2 in) tail 5–7 cm (2–3 in) arm/wing length[ 18] Habitat : Forest, shrubland, and caves[ 58]
LC
Unknown [ 58]
Guatemalan myotis
M. cobanensis Goodwin , 1955
Guatemala
Size : About 4 cm (2 in), plus about 4 cm (2 in) tail about 4 cm (2 in) arm/wing length[ 14] Habitat : Unknown[ 59]
DD
Unknown [ 59]
Hairy-faced bat
M. annectans Dobson , 1871
Southern and southeastern Asia
Size : 5–7 cm (2–3 in), plus 4–5 cm (2–2 in) tail 4–5 cm (2–2 in) arm/wing length[ 10] Habitat : Forest[ 60]
LC
Unknown [ 60]
Hairy-legged myotis
M. keaysi Allen , 1914
M. k. keaysi
M. k. pilosotibialis
Mexico, Central America, and northern and western South America
Size : 5–7 cm (2–3 in), plus 2–5 cm (1–2 in) tail Unknown arm/wing length[ 5] Habitat : Forest[ 61]
LC
Unknown [ 61]
Herman's myotis
M. hermani Thomas , 1923
Indonesia
Size : 5–6 cm (2–2 in), plus 3–6 cm (1–2 in) tail 4–6 cm (2–2 in) arm/wing length[ 7] Habitat : Forest[ 62]
DD
Unknown [ 62]
Himalayan whiskered bat
M. siligorensis Horsfield , 1855
M. s. alticraniatus
M. s. siligorensis
M. s. sowerbyi
M. s. thaianus
Central, eastern, and southeastern Asia
Size : Unknown length about 3 cm (1 in) arm/wing length[ 10] Habitat : Forest and caves[ 63]
LC
Unknown [ 63]
Hodgson's bat
M. formosus (Hodgson , 1835)
Central and eastern, and southeastern Asia
Size : 5–6 cm (2–2 in), plus 3–6 cm (1–2 in) tail 4–6 cm (2–2 in) arm/wing length[ 7] Habitat : Forest, shrubland, grassland, and caves[ 64]
NT
Unknown [ 64]
Horsfield's bat
M. horsfieldii Temminck , 1840
M. h. deignani
M. h. dryas
M. h. horsfieldii
M. h. jeannei
M. h. peshwa
Southern and southeastern Asia
Size : 4–6 cm (2–2 in), plus 3–5 cm (1–2 in) tail 3–5 cm (1–2 in) arm/wing length[ 10] Habitat : Forest and caves[ 65]
LC
Unknown [ 65]
Ikonnikov's bat
M. ikonnikovi Ogniov , 1912
Eastern Asia
Size : 3–6 cm (1–2 in), plus 2–5 cm (1–2 in) tail 3–4 cm (1–2 in) arm/wing length[ 7] Habitat : Forest, inland wetlands, and caves[ 66]
LC
Unknown [ 66]
Indiana bat
M. sodalis Miller , 1898
Eastern United States
Size : 4–5 cm (2–2 in), plus 2–5 cm (1–2 in) tail 3–4 cm (1–2 in) arm/wing length[ 5] Habitat : Forest and caves[ 67]
NT
Unknown [ 67]
Indochinese mouse-eared bat
M. indochinensis Son , Motokawa , Estók , Thong , Dang , Oshida , Csorba , Francis , Görföl , & Endō , 2013
Vietnam and China
Size : Unknown length 4–5 cm (2–2 in) arm/wing length[ 10] Habitat : Forest[ 68]
DD
Unknown [ 68]
Insular myotis
M. insularum Dobson , 1878
American Samoa
Size : About 4 cm (2 in), plus about 4 cm (2 in) tail about 4 cm (2 in) arm/wing length[ 10] Habitat : Unknown[ 69]
DD
Unknown [ 69]
Izecksohn's myotis
M. izecksohni Moratelli , Peracchi , Dias , & de Oliveira , 2011
Brazil and Argentina
Size : 4–7 cm (2–3 in), plus 3–5 cm (1–2 in) tail 3–4 cm (1–2 in) arm/wing length[ 14] Habitat : Forest[ 70]
DD
Unknown [ 70]
Kashmir cave bat
M. longipes Dobson , 1873
Southern Asia
Size : 3–5 cm (1–2 in), plus 3–5 cm (1–2 in) tail 3–4 cm (1–2 in) arm/wing length[ 10] Habitat : Forest and caves[ 71]
DD
Unknown [ 71]
Keen's myotis
M. keenii (Merriam , 1895)
Western Canada and United States
Size : 4–6 cm (2–2 in), plus 3–5 cm (1–2 in) tail 3–4 cm (1–2 in) arm/wing length[ 5] Habitat : Forest and rocky areas[ 72]
LC
Unknown [ 72]
Kei myotis
M. stalkeri Thomas , 1910
Indonesia
Size : 4–8 cm (2–3 in), plus 5–6 cm (2–2 in) tail 5–6 cm (2–2 in) arm/wing length[ 10] Habitat : Forest, inland wetlands, and caves[ 73]
LC
10,000–11,000 [ 73]
Kock's mouse-eared bat
M. dieteri Happold , 2005
Republic of the Congo
Size : About 5 cm (2 in), plus about 4 cm (2 in) tail about 4 cm (2 in) arm/wing length[ 7] Habitat : Forest and caves[ 74]
DD
Unknown [ 74]
Large myotis
M. chinensis Tomes , 1857
Eastern and southeastern Asia
Size : 9–10 cm (4–4 in), plus 5–6 cm (2–2 in) tail 6–7 cm (2–3 in) arm/wing length[ 18] Habitat : Forest, inland wetlands, and caves[ 75]
LC
Unknown [ 75]
Large-footed bat
M. adversus Horsfield , 1824
M. a. adversus
M. a. carimatae
M. a. orientis
M. a. taiwanensis
M. a. tanimbarensis
M. a. wetarensis
Southeastern Asia
Size : 4–6 cm (2–2 in), plus 3–5 cm (1–2 in) tail 3–5 cm (1–2 in) arm/wing length[ 10] Habitat : Forest, inland wetlands, and caves
LC
Unknown
Large-footed myotis
M. macropus (Gould , 1854)
Eastern Australia
Size : 3–5 cm (1–2 in), plus 3–5 cm (1–2 in) tail 3–5 cm (1–2 in) arm/wing length[ 18] Habitat : Inland wetlands and caves[ 77]
LC
Unknown [ 77]
LaVal's myotis
M. lavali Moratelli , Peracchi , Dias , & de Oliveira , 2011
Central and eastern South America
Size : 4–5 cm (2–2 in), plus 3–5 cm (1–2 in) tail 3–4 cm (1–2 in) arm/wing length[ 14] Habitat : Forest, savanna, and grassland[ 78]
LC
Unknown [ 78]
Lesser large-footed bat
M. hasseltii Temminck , 1840
M. h. abboti
M. h. continentis
M. h. hasseltii
M. h. macellus
Southern and southeastern Asia
Size : 4–6 cm (2–2 in), plus 3–5 cm (1–2 in) tail 3–5 cm (1–2 in) arm/wing length[ 18] Habitat : Forest and caves
LC
Unknown
Lesser mouse-eared bat
M. blythii Tomes , 1857
M. b. ancilla
M. b. blythii
M. b. lesviacus
M. b. omari
Europe and Asia
Size : 5–8 cm (2–3 in), plus about 6 cm (2 in) tail 5–7 cm (2–3 in) arm/wing length[ 18] Habitat : Shrubland, grassland, and caves[ 80]
LC
Unknown [ 80]
Little brown bat
M. lucifugus (Conte , 1831)
M. l. alascensis
M. l. carissima
M. l. lucifugus
M. l. pernox
M. l. relictus
United States and Canada
Size : 3–6 cm (1–2 in), plus 2–5 cm (1–2 in) tail 3–5 cm (1–2 in) arm/wing length[ 5] Habitat : Forest and caves[ 81]
EN
Unknown [ 81]
Long-eared myotis
M. evotis (H. Allen , 1864)
M. e. chrysonotus
M. e. evotis
M. e. jonesorum
M. e. micronyx
M. e. milleri
M. e. pacificus
Western North America
Size : 4–6 cm (2–2 in), plus 3–5 cm (1–2 in) tail 3–4 cm (1–2 in) arm/wing length[ 5] Habitat : Caves, rocky areas, and forest[ 82]
LC
Unknown [ 82]
Long-fingered bat
M. capaccinii Bonaparte , 1837
Southern Europe, northern Africa, and western Asia
Size : 4–6 cm (2–2 in), plus 3–5 cm (1–2 in) tail 3–5 cm (1–2 in) arm/wing length[ 10] Habitat : Shrubland, inland wetlands, and caves[ 83]
VU
Unknown [ 83]
Long-legged myotis
M. volans H. Allen , 1866
M. v. amotus
M. v. interior
M. v. longicrus
M. v. volans
Western North America
Size : 4–7 cm (2–3 in), plus 4–5 cm (2–2 in) tail 3–5 cm (1–2 in) arm/wing length[ 5] Habitat : Forest, rocky areas, caves, and desert[ 84]
LC
Unknown [ 84]
Long-tailed myotis
M. longicaudatus Ogniov , 1927
M. l. eniseensis
M. l. kaguyae
M. l. longicaudatus
Eastern Asia (excluding red)
Size : 4–6 cm (2–2 in), plus 3–5 cm (1–2 in) tail 3–5 cm (1–2 in) arm/wing length[ 18] Habitat : Forest, inland wetlands, and caves[ 85]
LC
Unknown [ 85]
Long-toed myotis
M. secundus Ruedi , Csorba , Lin , & Chou , 2015
Taiwan
Size : 3–5 cm (1–2 in), plus 3–5 cm (1–2 in) tail 3–4 cm (1–2 in) arm/wing length[ 10] Habitat : Forest[ 86]
LC
Unknown [ 86]
Malagasy mouse-eared bat
M. goudoti Smith , 1834
Madagascar
Size : Unknown length 3–4 cm (1–2 in) arm/wing length[ 7] Habitat : Forest, rocky areas, and caves[ 87]
LC
Unknown [ 87]
Malaysian whiskered myotis
M. federatus Thomas , 1916
Malaysia (in blue)
Size : About 5 cm (2 in), plus about 4 cm (2 in) tail 3–5 cm (1–2 in) arm/wing length[ 10] Habitat : Forest[ 88]
DD
Unknown [ 88]
Maluku myotis
M. moluccarum Thomas , 1915
M. m. moluccarum
M. m. richardsi
M. m. solomonis
Indonesia
Size : 4–6 cm (2–2 in), plus 2–5 cm (1–2 in) tail 3–5 cm (1–2 in) arm/wing length[ 10] Habitat : Forest and inland wetlands[ 89]
LC
Unknown [ 89]
Mandelli's mouse-eared bat
M. sicarius Thomas , 1915
India and Nepal
Size : 5–6 cm (2–2 in), plus 4–6 cm (2–2 in) tail 4–6 cm (2–2 in) arm/wing length[ 18] Habitat : Forest and caves[ 90]
VU
Unknown [ 90]
Montane myotis
M. oxyotus Peters , 1867
M. o. gardneri
M. o. oxyotus
Northern and western South America
Size : 4–6 cm (2–2 in), plus 4–5 cm (2–2 in) tail 3–5 cm (1–2 in) arm/wing length[ 14] Habitat : Forest[ 91]
LC
Unknown [ 91]
Morris's bat
M. morrisi Hill , 1971
Ethiopia and Nigeria
Size : About 5 cm (2 in), plus about 5 cm (2 in) tail 4–5 cm (2–2 in) arm/wing length[ 7] Habitat : Savanna and caves[ 92]
DD
Unknown [ 92]
Natterer's bat
M. nattereri (Kuhl , 1817)
M. n. nattereri
M. n. tschuliensis
Europe and western Asia
Size : 4–5 cm (2–2 in), plus 3–5 cm (1–2 in) tail 3–5 cm (1–2 in) arm/wing length[ 18] Habitat : Forest, shrubland, grassland, inland wetlands, and caves[ 93]
LC
Unknown [ 93]
Nepal myotis
M. nipalensis Dobson , 1871
M. n. nipalensis
M. n. przewalskii
M. n. transcaspicus
Central and western Asia
Size : About 4 cm (2 in), plus about 33–5 cm (1–2 in) tail about 3 cm (1 in) arm/wing length[ 7] Habitat : Forest, shrubland, grassland, caves, and desert[ 94]
LC
Unknown [ 94]
Nimba myotis
M. nimbaensis Simmons , Flanders , Fils , Parker , Suter , Bamba , Douno , Keita , Morales , & Frick , 2021
Guinea
Size : 6–8 cm (2–3 in), plus 4–5 cm (2–2 in) tail 5–6 cm (2–2 in) arm/wing length[ 7] Habitat : Forest, grassland, inland wetlands, caves, and shrubland[ 95]
CR
Unknown [ 95]
Northern long-eared bat
M. septentrionalis Trouessart , 1897
Canada and eastern United States
Size : 4–5 cm (2–2 in), plus 3–5 cm (1–2 in) tail 3–4 cm (1–2 in) arm/wing length[ 5] Habitat : Forest and caves[ 96]
NT
Unknown [ 96]
Orange-fingered myotis
M. rufopictus (Waterhouse , 1845)
Indonesia
Size : About 6 cm (2 in), plus 4–6 cm (2–2 in) tail 4–6 cm (2–2 in) arm/wing length[ 7] Habitat : Forest[ 97]
DD
Unknown [ 97]
Pallid large-footed myotis
M. macrotarsus Waterhouse , 1845
M. m. macrotarsus
M. m. saba
Indonesia
Size : About 6 cm (2 in), plus 4–6 cm (2–2 in) tail 4–5 cm (2–2 in) arm/wing length[ 10] Habitat : Caves
LC
Unknown
Peninsular myotis
M. peninsularis Miller , 1898
Southern Baja California peninsula
Size : 4–6 cm (2–2 in), plus 3–6 cm (1–2 in) tail 3–5 cm (1–2 in) arm/wing length[ 14] Habitat : Forest and caves[ 99]
EN
Unknown [ 99]
Peters's myotis
M. ater (Peters , 1866)
Southeastern Asia
Size : 4–6 cm (2–2 in), plus 3–5 cm (1–2 in) tail 3–5 cm (1–2 in) arm/wing length[ 10] Habitat : Forest
LC
Unknown
Peyton's myotis
M. peytoni Wroughton & Ryley , 1913
Southern India (in yellow)
Size : 5–7 cm (2–3 in), plus 4–5 cm (2–2 in) tail 4–5 cm (2–2 in) arm/wing length[ 10] Habitat : Forest[ 101]
DD
Unknown [ 101]
Pond bat
M. dasycneme (Boie , 1825)
Europe and northern Asia
Size : 5–7 cm (2–3 in), plus 4–6 cm (2–2 in) tail 4–5 cm (2–2 in) arm/wing length[ 7] Habitat : Forest, inland wetlands, and caves[ 102]
NT
Unknown [ 102]
Red myotis
M. ruber Geoffroy , 1806
Southeastern South America
Size : 4–5 cm (2–2 in), plus 3–5 cm (1–2 in) tail 3–5 cm (1–2 in) arm/wing length[ 5] Habitat : Forest[ 103]
NT
Unknown [ 103]
Reddish myotis
M. soror Ruedi , Csorba , Lin , & Chou , 2015
Taiwan
Size : About 5 cm (2 in), plus about 4 cm (2 in) tail about 4 cm (2 in) arm/wing length[ 18] Habitat : Forest[ 104]
DD
Unknown [ 104]
Reddish-black myotis
M. rufoniger (Tomes , 1858)
Eastern Asia
Size : 5–8 cm (2–3 in), plus 4–6 cm (2–2 in) tail 4–6 cm (2–2 in) arm/wing length[ 7] Habitat : Forest and caves[ 105]
LC
Unknown [ 105]
Rickett's big-footed bat
M. pilosus Peters , 1869
Eastern Asia
Size : 5–7 cm (2–3 in), plus 3–6 cm (1–2 in) tail 5–7 cm (2–3 in) arm/wing length[ 18] Habitat : Forest and inland wetlands
VU
Unknown
Ridley's bat
M. ridleyi Thomas , 1898
Southeastern Asia
Size : 3–5 cm (1–2 in), plus 2–4 cm (1–2 in) tail 2–4 cm (1–2 in) arm/wing length[ 10] Habitat : Caves, inland wetlands, and forest[ 107]
NT
Unknown [ 107]
Riparian myotis
M. riparius Handley , 1960
Central America and South America
Size : 4–5 cm (2–2 in), plus 2–5 cm (1–2 in) tail 3–4 cm (1–2 in) arm/wing length[ 14] Habitat : Forest[ 108]
LC
Unknown [ 108]
Rufous mouse-eared bat
M. bocagii Peters , 1870
M. b. bocagii
M. b. cupreolus
M. b. dogalensis
Scattered sub-Saharan Africa and Yemen
Size : 5–6 cm (2–2 in), plus 3–5 cm (1–2 in) tail 3–5 cm (1–2 in) arm/wing length[ 7] Habitat : Forest and savanna[ 109]
LC
Unknown [ 109]
Schaub's myotis
M. schaubi Kormos , 1934
M. s. araxenus
M. s. schaubi
Armenia and Iran
Size : 4–6 cm (2–2 in), plus 4–5 cm (2–2 in) tail 3–5 cm (1–2 in) arm/wing length[ 18] Habitat : Forest, shrubland, and caves[ 110]
DD
Unknown [ 110]
Schwartz's myotis
M. martiniquensis LaVal , 1973
Barbados and Martinique in the Caribbean
Size : Unknown length 3–4 cm (1–2 in) arm/wing length[ 14] Habitat : Caves[ 111]
NT
Unknown [ 111]
Scott's mouse-eared bat
M. scotti Thomas , 1927
Ethiopia
Size : 4–6 cm (2–2 in), plus 4–5 cm (2–2 in) tail 3–5 cm (1–2 in) arm/wing length[ 7] Habitat : Forest and shrubland[ 112]
VU
Unknown [ 112]
Siberian bat
M. sibiricus (Kastschenko , 1905)
Northeastern Asia (in green)
Size : 3–6 cm (1–2 in), plus 3–5 cm (1–2 in) tail 3–4 cm (1–2 in) arm/wing length[ 7] Habitat : Forest, savanna, rocky areas, and caves[ 113]
LC
Unknown [ 113]
Silver-tipped myotis
M. albescens Geoffroy , 1806
Central America and South America
Size : 4–5 cm (2–2 in), plus 3–5 cm (1–2 in) tail 3–4 cm (1–2 in) arm/wing length[ 14] Habitat : Forest[ 114]
LC
Unknown [ 114]
Singapore whiskered bat
M. oreias (Temminck , 1840)
Singapore
Size : Unknown[ 115] Habitat : Unknown[ 115]
DD
Unknown [ 115]
Southeastern myotis
M. austroriparius (Rhoads , 1897)
Southeastern United States
Size : 4–6 cm (2–2 in), plus 2–5 cm (1–2 in) tail 3–4 cm (1–2 in) arm/wing length[ 14] Habitat : Forest and caves[ 116]
LC
Unknown [ 116]
Southern myotis
M. aelleni Baud , 1979
Argentina
Size : 4–5 cm (2–2 in), plus 2–4 cm (1–2 in) tail 3–5 cm (1–2 in) arm/wing length[ 14] Habitat : Unknown[ 117]
DD
Unknown [ 117]
Southwestern myotis
M. auriculus Baker & Stains , 1955
M. a. apache
M. a. auriculus
Southern United States and Mexico
Size : 5–6 cm (2–2 in), plus 3–5 cm (1–2 in) tail 3–4 cm (1–2 in) arm/wing length[ 5] Habitat : Forest, caves, and desert[ 118]
LC
Unknown [ 118]
Szechwan myotis
M. altarium Thomas , 1911
China and Thailand
Size : 4–6 cm (2–2 in), plus 3–5 cm (1–2 in) tail 4–5 cm (2–2 in) arm/wing length[ 7] Habitat : Caves[ 119]
LC
Unknown [ 119]
Thick-thumbed myotis
M. rosseti Oei , 1951
Southeastern Asia
Size : 3–5 cm (1–2 in), plus 3–5 cm (1–2 in) tail 2–4 cm (1–2 in) arm/wing length[ 10] Habitat : Forest[ 120]
LC
Unknown [ 120]
Velvety myotis
M. simus Thomas , 1901
Western, central, and eastern South America
Size : 5–6 cm (2–2 in), plus 3–4 cm (1–2 in) tail 3–4 cm (1–2 in) arm/wing length[ 14] Habitat : Forest[ 121]
DD
Unknown [ 121]
Wall-roosting mouse-eared bat
M. muricola (Gray , 1846)
M. m. browni
M. m. caliginosus
M. m. herrei
M. m. moupinensis
M. m. muricola
M. m. niasensis
M. m. patriciae
Southeastern Asia
Size : 4–6 cm (2–2 in), plus 3–5 cm (1–2 in) tail 3–4 cm (1–2 in) arm/wing length[ 10] Habitat : Forest and caves[ 122]
LC
Unknown [ 122]
Weber's myotis
M. weberi (Jentink , 1890)
Sulawesi island in Indonesia
Size : About 6 cm (2 in), plus about 4 cm (2 in) tail 4–6 cm (2–2 in) arm/wing length[ 7] Habitat : Forest[ 123]
DD
Unknown [ 123]
Welwitsch's bat
M. welwitschii (Gray , 1866)
Western, eastern, and southern Africa
Size : 5–7 cm (2–3 in), plus 5–7 cm (2–3 in) tail 5–6 cm (2–2 in) arm/wing length[ 7] Habitat : Forest, savanna, shrubland, and grassland[ 124]
LC
Unknown [ 124]
Western small-footed bat
M. ciliolabrum (Merriam , 1842)
Western North America
Size : 3–5 cm (1–2 in), plus 2–5 cm (1–2 in) tail 3–4 cm (1–2 in) arm/wing length[ 5] Habitat : Forest, rocky areas, caves, and desert
LC
Unknown
Whiskered bat
M. mystacinus (Kuhl , 1817)
M. m. caucasicus
M. m. mystacinus
M. m. occidentalis
Europe, northern Africa, and western Asia
Size : 3–5 cm (1–2 in), plus 3–5 cm (1–2 in) tail 3–4 cm (1–2 in) arm/wing length[ 7] Habitat : Desert, caves, grassland, shrubland, and forest[ 126]
LC
Unknown [ 126]
Yanbaru whiskered bat
M. yanbarensis Maeda & Matsumura , 1998
Ryukyu Islands in Japan
Size : 3–5 cm (1–2 in), plus 3–5 cm (1–2 in) tail 3–4 cm (1–2 in) arm/wing length[ 10] Habitat : Forest[ 127]
CR
Unknown [ 127]
Yellowish myotis
M. levis Geoffroy , 1806
Southeastern South America (in red)
Size : 3–6 cm (1–2 in), plus 4–5 cm (2–2 in) tail 3–5 cm (1–2 in) arm/wing length[ 14] Habitat : Forest[ 128]
LC
Unknown [ 128]
Yuma myotis
M. yumanensis H. Allen , 1864
M. y. lambi
M. y. lutosus
M. y. oxalis
M. y. saturatus
M. y. sociabilis
M. y. yumanensis
Western North America
Size : 4–5 cm (2–2 in), plus 2–4 cm (1–2 in) tail 3–4 cm (1–2 in) arm/wing length[ 14] Habitat : Forest and caves[ 129]
LC
Unknown [ 129]
Genus Submyotodon – Ziegler , 2003 – one species
Common name
Scientific name and subspecies
Range
Size and ecology
IUCN status and estimated population
Taiwan broad-muzzled bat
S. latirostris (Kishida , 1932)
Taiwan
Size : 3–4 cm (1–2 in), plus 3–4 cm (1–2 in) tail 3–4 cm (1–2 in) arm/wing length[ 5] Habitat : Forest[ 130]
LC
Unknown [ 130]
References
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^ "Fossilworks: Myotis" . Paleobiology Database . University of Wisconsin–Madison . Archived from the original on September 22, 2022. Retrieved November 2, 2024 .
^ Wilson, Reeder , pp. 499–518
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^ a b c d e f g h i j k l m n o p q r s t u v w x y Chernasky; Motis; Burgin , pp. 560–561
^ a b Hutson, A. M.; Paunović, M. (2016). "Myotis alcathoe " . IUCN Red List of Threatened Species . 2016 : e.T136680A518740. doi :10.2305/IUCN.UK.2016-2.RLTS.T136680A518740.en .
^ a b Jacobs, D. (2019). "Myotis anjouanensis " . IUCN Red List of Threatened Species . 2019 : e.T44863A22073545. doi :10.2305/IUCN.UK.2019-3.RLTS.T44863A22073545.en .
^ a b c d e f g h i j k l m n o p q r s t u v w x y z Chernasky; Motis; Burgin , pp. 562–563
^ a b Srinivasulu, C.; Srinivasulu, B. (2019). "Myotis annatessae " . IUCN Red List of Threatened Species . 2019 : e.T85342605A85342608. doi :10.2305/IUCN.UK.2019-3.RLTS.T85342605A85342608.en .
^ a b Kruskop, S. V. (2016). "Myotis annamiticus " . IUCN Red List of Threatened Species . 2016 : e.T136279A22006224. doi :10.2305/IUCN.UK.2016-2.RLTS.T136279A22006224.en .
^ a b Solari, S. (2018). "Myotis occultus " . IUCN Red List of Threatened Species . 2018 : e.T136650A21990499. doi :10.2305/IUCN.UK.2018-2.RLTS.T136650A21990499.en .
^ a b c d e f g h i j k l m n o p q r s t u v w x Chernasky; Motis; Burgin , pp. 558–559
^ a b Vargas-Rodríguez, R.; Peñaranda, D.; Ugarte Nuñez, J.; Rodríguez-San Pedro, A.; Ossa Gomez, G.; Gatica Castro, A. (2016). "Myotis atacamensis " . IUCN Red List of Threatened Species . 2016 : e.T14143A22050638. doi :10.2305/IUCN.UK.2016-1.RLTS.T14143A22050638.en .
^ a b c Reardon, T. B.; Lumsden, L. F. (2020). "Myotis australis " . IUCN Red List of Threatened Species . 2020 : e.T14146A22060248. doi :10.2305/IUCN.UK.2020-3.RLTS.T14146A22060248.en .
^ a b Larsen, R. (2016). "Myotis nyctor " . IUCN Red List of Threatened Species . 2016 : e.T76435059A76435083. doi :10.2305/IUCN.UK.2016-1.RLTS.T76435059A76435083.en .
^ a b c d e f g h i j k l m n o p q r s t u v w Chernasky; Motis; Burgin , pp. 564–565
^ a b Paunović, M. (2016). "Myotis bechsteinii " . IUCN Red List of Threatened Species . 2016 : e.T14123A22053752. doi :10.2305/IUCN.UK.2016-2.RLTS.T14123A22053752.en .
^ a b Feng, J.; Jiang, T. L. (2019). "Myotis pequinius " . IUCN Red List of Threatened Species . 2019 : e.T14190A22066613. doi :10.2305/IUCN.UK.2019-3.RLTS.T14190A22066613.en .
^ a b Solari, S. (2019). "Myotis nigricans " . IUCN Red List of Threatened Species . 2019 : e.T14185A22066939. doi :10.2305/IUCN.UK.2019-2.RLTS.T14185A22066939.en .
^ a b Srinivasulu, B.; Srinivasulu, C. (2019). "Myotis bucharensis " . IUCN Red List of Threatened Species . 2019 : e.T136219A22011494. doi :10.2305/IUCN.UK.2019-3.RLTS.T136219A22011494.en .
^ a b Görföl, T.; Csorba, G. (2017). "Myotis borneoensis " . IUCN Red List of Threatened Species . 2017 : e.T85568289A85568292. doi :10.2305/IUCN.UK.2017-2.RLTS.T85568289A85568292.en .
^ a b Gazaryan, S.; Kruskop, S. V.; Godlevska, L. (2021) [errata version of 2020 assessment]. "Myotis brandtii " . IUCN Red List of Threatened Species . 2020 : e.T85566997A195857637. doi :10.2305/IUCN.UK.2020-2.RLTS.T85566997A195857637.en .
^ a b Görföl, T.; Tu, V.; Csorba, G.; Francis, C. M.; Hutson, A. M.; Bates, P. J. J.; Bumrungsri, S.; Molur, S.; Srinivasulu, C. (2020). "Myotis montivagus " . IUCN Red List of Threatened Species . 2020 : e.T85567622A22065126. doi :10.2305/IUCN.UK.2020-2.RLTS.T85567622A22065126.en .
^ a b Arroyo-Cabrales, J.; Perez, S. (2017). "Myotis californicus " . IUCN Red List of Threatened Species . 2017 : e.T14150A22061366. doi :10.2305/IUCN.UK.2017-2.RLTS.T14150A22061366.en .
^ a b Monadjem, A.; Jacobs, D. (2017). "Myotis tricolor " . IUCN Red List of Threatened Species . 2017 : e.T14207A22063832. doi :10.2305/IUCN.UK.2017-2.RLTS.T14207A22063832.en .
^ a b Solari, S. (2019). "Myotis velifer " . IUCN Red List of Threatened Species . 2019 : e.T14208A22063586. doi :10.2305/IUCN.UK.2019-1.RLTS.T14208A22063586.en .
^ a b Barquez, R.; Diaz, M. (2016). "Myotis chiloensis " . IUCN Red List of Threatened Species . 2016 : e.T14151A22061103. doi :10.2305/IUCN.UK.2016-3.RLTS.T14151A22061103.en .
^ a b Feng, J.; Jiang, T. L. (2019). "Myotis laniger " . IUCN Red List of Threatened Species . 2019 : e.T136429A21984685. doi :10.2305/IUCN.UK.2019-3.RLTS.T136429A21984685.en .
^ a b Perez, S.; de Grammont, P. C.; Cuarón, A. D. (2017). "Myotis fortidens " . IUCN Red List of Threatened Species . 2017 : e.T14161A22056846. doi :10.2305/IUCN.UK.2017-2.RLTS.T14161A22056846.en .
^ a b Russo, D.; Cistrone, L. (2024) [errata version of 2023 assessment]. "Myotis crypticus " . IUCN Red List of Threatened Species . 2023 : e.T215154989A254355251.
^ a b c Csorba, G.; Thapa, S. (2016). "Myotis csorbai " . IUCN Red List of Threatened Species . 2016 : e.T29420A22070788. doi :10.2305/IUCN.UK.2016-2.RLTS.T29420A22070788.en .
^ a b Solari, S. (2016). "Myotis nesopolus " . IUCN Red List of Threatened Species . 2016 : e.T14184A22065759. doi :10.2305/IUCN.UK.2016-1.RLTS.T14184A22065759.en .
^ a b Kruskop, S. V.; Godlevska, L.; Bücs, S.; Çoraman, E.; Gazaryan, S. (2021) [errata version of 2020 assessment]. "Myotis daubentonii " . IUCN Red List of Threatened Species . 2020 : e.T85342710A195858793. doi :10.2305/IUCN.UK.2020-2.RLTS.T85342710A195858793.en .
^ a b Jiang, T. L.; Feng, J. (2019). "Myotis davidii " . IUCN Red List of Threatened Species . 2019 : e.T136250A22003049. doi :10.2305/IUCN.UK.2019-3.RLTS.T136250A22003049.en .
^ a b Solari, S. (2017). "Myotis diminutus " . IUCN Red List of Threatened Species . 2017 : e.T88151417A88151431. doi :10.2305/IUCN.UK.2017-2.RLTS.T88151417A88151431.en .
^ a b Barquez, R.; Diaz, M. (2016). "Myotis dinellii " . IUCN Red List of Threatened Species . 2016 : e.T136204A22009702. doi :10.2305/IUCN.UK.2016-3.RLTS.T136204A22009702.en .
^ a b Larsen, R. (2016). "Myotis dominicensis " . IUCN Red List of Threatened Species . 2016 : e.T14155A22057933. doi :10.2305/IUCN.UK.2016-1.RLTS.T14155A22057933.en .
^ a b Fukui, D.; Sun, K.; Kruskop, S. V. (2019). "Myotis macrodactylus " . IUCN Red List of Threatened Species . 2019 : e.T14177A22065868. doi :10.2305/IUCN.UK.2019-3.RLTS.T14177A22065868.en .
^ a b Fukui, D.; Sano, A. (2020). "Myotis petax " . IUCN Red List of Threatened Species . 2020 : e.T85342726A85342734. doi :10.2305/IUCN.UK.2020-2.RLTS.T85342726A85342734.en .
^ a b Miller, B.; Rodriguez, B. (2017) [errata version of 2016 assessment]. "Myotis elegans " . IUCN Red List of Threatened Species . 2016 : e.T14156A115121563. doi :10.2305/IUCN.UK.2016-3.RLTS.T14156A22057814.en .
^ a b Russo, D.; Cistrone, L. (2023). "Myotis escalerai " . IUCN Red List of Threatened Species . 2023 : e.T85733126A211003991. doi :10.2305/IUCN.UK.2023-1.RLTS.T85733126A211003991.en .
^ a b Fukui, D.; Sano, A.; Kruskop, S. V. (2019). "Myotis bombinus " . IUCN Red List of Threatened Species . 2019 : e.T14149A22061650. doi :10.2305/IUCN.UK.2019-3.RLTS.T14149A22061650.en .
^ a b Juste, J.; Paunović, M. (2016). "Myotis punicus " . IUCN Red List of Threatened Species . 2016 : e.T44864A22073410. doi :10.2305/IUCN.UK.2016-2.RLTS.T44864A22073410.en .
^ a b Arroyo-Cabrales, J.; Ospina-Garces, S. (2016). "Myotis vivesi " . IUCN Red List of Threatened Species . 2016 : e.T14209A22069146. doi :10.2305/IUCN.UK.2016-1.RLTS.T14209A22069146.en .
^ a b Arroyo-Cabrales, J.; Ospina-Garces, S. (2016). "Myotis findleyi " . IUCN Red List of Threatened Species . 2016 : e.T14159A22058800. doi :10.2305/IUCN.UK.2016-1.RLTS.T14159A22058800.en .
^ a b Arroyo-Cabrales, J.; Ospina-Garces, S. (2016). "Myotis planiceps " . IUCN Red List of Threatened Species . 2016 : e.T14191A22066742. doi :10.2305/IUCN.UK.2016-1.RLTS.T14191A22066742.en .
^ a b Vincenot, C. E.; Preble, J. H.; Huang, J. C. -C.; Collazo, A. M.; Kamal, A. (2021). "Myotis frater " . IUCN Red List of Threatened Species . 2021 : e.T85566806A22056940. doi :10.2305/IUCN.UK.2021-2.RLTS.T85566806A22056940.en .
^ a b Arroyo-Cabrales, J.; de Grammont, P. C. (2017). "Myotis thysanodes " . IUCN Red List of Threatened Species . 2017 : e.T14206A22063246. doi :10.2305/IUCN.UK.2017-2.RLTS.T14206A22063246.en .
^ a b Fukui, D.; Sano, A. (2021) [errata version of 2019 assessment]. "Myotis pruinosus " . IUCN Red List of Threatened Species . 2019 : e.T14192A209551299. doi :10.2305/IUCN.UK.2019-3.RLTS.T14192A209551299.en .
^ a b Piraccini, R. (2016). "Myotis emarginatus " . IUCN Red List of Threatened Species . 2016 : e.T14129A22051191. doi :10.2305/IUCN.UK.2016-2.RLTS.T14129A22051191.en .
^ a b Waldien, D. L.; Santiago, K.; Wortham, G.; Stronsick, S. (2021). "Myotis gomantongensis " . IUCN Red List of Threatened Species . 2021 : e.T40035A22060096. doi :10.2305/IUCN.UK.2021-3.RLTS.T40035A22060096.en .
^ a b Solari, S. (2018). "Myotis grisescens " . IUCN Red List of Threatened Species . 2018 : e.T14132A22051652. doi :10.2305/IUCN.UK.2018-2.RLTS.T14132A22051652.en .
^ a b Coroiu, I.; Juste, J.; Paunović, M. (2016). "Myotis myotis " . IUCN Red List of Threatened Species . 2016 : e.T14133A22051759. doi :10.2305/IUCN.UK.2016-2.RLTS.T14133A22051759.en .
^ a b Cajas C., J.; Miller, B. (2016). "Myotis cobanensis " . IUCN Red List of Threatened Species . 2016 : e.T14154A22058031. doi :10.2305/IUCN.UK.2016-2.RLTS.T14154A22058031.en .
^ a b Görföl, T.; Csorba, G.; Bates, P. J. J.; Francis, C. M.; Molur, S.; Srinivasulu, C. (2020). "Myotis annectans " . IUCN Red List of Threatened Species . 2020 : e.T14142A22050272. doi :10.2305/IUCN.UK.2020-2.RLTS.T14142A22050272.en .
^ a b Barquez, R.; Diaz, M. (2016). "Myotis keaysi " . IUCN Red List of Threatened Species . 2016 : e.T14170A22056048. doi :10.2305/IUCN.UK.2016-3.RLTS.T14170A22056048.en .
^ a b Csorba, G.; Hutson, A. M.; Kingston, T.; Bumrungsri, S.; Francis, C. M. (2016). "Myotis hermani " . IUCN Red List of Threatened Species . 2016 : e.T14165A22057251. doi :10.2305/IUCN.UK.2016-2.RLTS.T14165A22057251.en .
^ a b Santiago, K.; Wortham, G.; Waldien, D. L. (2021). "Myotis siligorensis " . IUCN Red List of Threatened Species . 2021 : e.T14203A22064839. doi :10.2305/IUCN.UK.2021-3.RLTS.T14203A22064839.en .
^ a b Huang, J. C. -C.; Csorba, G.; Chang, H.-C.; Ho, Y.-Y. (2020). "Myotis formosus " . IUCN Red List of Threatened Species . 2020 : e.T85736120A95642290. doi :10.2305/IUCN.UK.2020-2.RLTS.T85736120A95642290.en .
^ a b Phelps, K.; Rosell-Ambal, R. G. B.; Tabaranza, B.; Heaney, L.; Gonzalez, J. C.; Molur, S.; Srinivasulu, C. (2019). "Myotis horsfieldii " . IUCN Red List of Threatened Species . 2019 : e.T14166A22057415. doi :10.2305/IUCN.UK.2019-3.RLTS.T14166A22057415.en .
^ a b Zhigalin, A.; Stubbe, M.; Ariunbold, J.; Buuveibaatar, V.; Dorjderem, S.; Monkhzul, T.; Otgonbaatar, M.; Tsogbadrakh, M. (2020). "Myotis ikonnikovi " . IUCN Red List of Threatened Species . 2020 : e.T14168A22057122. doi :10.2305/IUCN.UK.2020-3.RLTS.T14168A22057122.en .
^ a b Arroyo-Cabrales, J.; Ospina-Garces, S. (2016). "Myotis sodalis " . IUCN Red List of Threatened Species . 2016 : e.T14136A22053184. doi :10.2305/IUCN.UK.2016-1.RLTS.T14136A22053184.en .
^ a b Son, N.; Görföl, T.; Csorba, G. (2019). "Myotis indochinensis " . IUCN Red List of Threatened Species . 2019 : e.T85342688A85342691. doi :10.2305/IUCN.UK.2019-3.RLTS.T85342688A85342691.en .
^ a b Helgen, K.; Bonaccorso, F. J. (2020). "Myotis insularum " . IUCN Red List of Threatened Species . 2020 : e.T14169A22055968. doi :10.2305/IUCN.UK.2020-2.RLTS.T14169A22055968.en .
^ a b Solari, S. (2017). "Myotis izecksohni " . IUCN Red List of Threatened Species . 2017 : e.T88151563A88151572. doi :10.2305/IUCN.UK.2017-2.RLTS.T88151563A88151572.en .
^ a b Kruskop, S. V. (2016). "Myotis longipes " . IUCN Red List of Threatened Species . 2016 : e.T14175A22056206. doi :10.2305/IUCN.UK.2016-2.RLTS.T14175A22056206.en .
^ a b Arroyo-Cabrales, J.; Álvarez-Castañeda, S. T. (2017). "Myotis keenii " . IUCN Red List of Threatened Species . 2017 : e.T14171A22055579. doi :10.2305/IUCN.UK.2017-2.RLTS.T14171A22055579.en .
^ a b Bouillard, N. (2021). "Myotis stalkeri " . IUCN Red List of Threatened Species . 2021 : e.T14205A22063416. doi :10.2305/IUCN.UK.2021-3.RLTS.T14205A22063416.en .
^ a b Happold. M. (2019). "Myotis dieteri " . IUCN Red List of Threatened Species . 2019 : e.T136678A22038629. doi :10.2305/IUCN.UK.2019-3.RLTS.T136678A22038629.en .
^ a b Jiang, T. L.; Feng, J. (2019). "Myotis chinensis " . IUCN Red List of Threatened Species . 2019 : e.T14152A22060946. doi :10.2305/IUCN.UK.2019-3.RLTS.T14152A22060946.en .
^ a b Gorecki, V.; Pennay, M. (2021). "Myotis macropus " . IUCN Red List of Threatened Species . 2021 : e.T136697A22039960. doi :10.2305/IUCN.UK.2021-1.RLTS.T136697A22039960.en .
^ a b Solari, S. (2017). "Myotis lavali " . IUCN Red List of Threatened Species . 2017 : e.T88151601A88151604. doi :10.2305/IUCN.UK.2017-2.RLTS.T88151601A88151604.en .
^ a b Juste, J.; Paunović, M. (2016). "Myotis blythii " . IUCN Red List of Threatened Species . 2016 : e.T14124A22053297. doi :10.2305/IUCN.UK.2016-2.RLTS.T14124A22053297.en .
^ a b Solari, S. (2021) [amended version of 2018 assessment]. "Myotis lucifugus " . IUCN Red List of Threatened Species . 2021 : e.T14176A208031565. doi :10.2305/IUCN.UK.2021-3.RLTS.T14176A208031565.en .
^ a b Arroyo-Cabrales, J.; Álvarez-Castañeda, S. T. (2017). "Myotis evotis " . IUCN Red List of Threatened Species . 2017 : e.T14157A22059133. doi :10.2305/IUCN.UK.2017-2.RLTS.T14157A22059133.en .
^ a b Paunović, M. (2016). "Myotis capaccinii " . IUCN Red List of Threatened Species . 2016 : e.T14126A22054131. doi :10.2305/IUCN.UK.2016-2.RLTS.T14126A22054131.en .
^ a b Solari, S. (2019). "Myotis volans " . IUCN Red List of Threatened Species . 2019 : e.T14210A22069325. doi :10.2305/IUCN.UK.2019-1.RLTS.T14210A22069325.en .
^ a b Vincenot, C. E.; Preble, J. H.; Collazo, A. M. (2021). "Myotis longicaudatus " . IUCN Red List of Threatened Species . 2021 : e.T85566977A85566980. doi :10.2305/IUCN.UK.2021-2.RLTS.T85566977A85566980.en .
^ a b Ruedi, M.; Csorba, G.; Liang-Kong, L.; Cheng-Han, C. (2017). "Myotis secundus " . IUCN Red List of Threatened Species . 2017 : e.T85342651A85342654. doi :10.2305/IUCN.UK.2017-2.RLTS.T85342651A85342654.en .
^ a b Monadjem, A.; Razafimanahaka, J.; Ranivo, J.; Kofoky, A.; Hutson, A. M.; Cardiff, S. G.; Andriafidison, D.; Goodman, S.; Jenkins, R. K. B.; Racey, P. A.; Ratrimomanarivo, F. H. (2017). "Myotis goudoti " . IUCN Red List of Threatened Species . 2017 : e.T14163A22056541. doi :10.2305/IUCN.UK.2017-2.RLTS.T14163A22056541.en .
^ a b Görföl, T.; Csorba, G. (2017). "Myotis federatus " . IUCN Red List of Threatened Species . 2017 : e.T85568302A85568305. doi :10.2305/IUCN.UK.2017-2.RLTS.T85568302A85568305.en .
^ a b Armstrong, K. N. (2021). "Myotis moluccarum " . IUCN Red List of Threatened Species . 2021 : e.T136770A22033795. doi :10.2305/IUCN.UK.2021-3.RLTS.T136770A22033795.en .
^ a b Srinivasulu, B.; Srinivasulu, C. (2019). "Myotis sicarius " . IUCN Red List of Threatened Species . 2019 : e.T14202A22063965. doi :10.2305/IUCN.UK.2019-3.RLTS.T14202A22063965.en .
^ a b Solari, S. (2018). "Myotis oxyotus " . IUCN Red List of Threatened Species . 2018 : e.T14187A22067211. doi :10.2305/IUCN.UK.2018-2.RLTS.T14187A22067211.en .
^ a b Jacobs, D.; Cotterill, F. P. D.; Taylor, P. J. (2019). "Myotis morrisi " . IUCN Red List of Threatened Species . 2019 : e.T14182A22065314. doi :10.2305/IUCN.UK.2019-3.RLTS.T14182A22065314.en .
^ a b Russo, D.; Cistrone, L. (2023). "Myotis nattereri " . IUCN Red List of Threatened Species . 2023 : e.T215492021A211005466. doi :10.2305/IUCN.UK.2023-1.RLTS.T215492021A211005466.en .
^ a b Srinivasulu, B.; Srinivasulu, C. (2019). "Myotis nipalensis " . IUCN Red List of Threatened Species . 2019 : e.T136495A21976309. doi :10.2305/IUCN.UK.2019-3.RLTS.T136495A21976309.en .
^ a b Bakwo Fils, E. M.; Flanders, J.; Frick, W. F.; Simmons, N. (2022). "Myotis nimbaensis " . IUCN Red List of Threatened Species . 2022 : e.T216617275A216617367. doi :10.2305/IUCN.UK.2022-2.RLTS.T216617275A216617367.en .
^ a b Solari, S. (2018). "Myotis septentrionalis " . IUCN Red List of Threatened Species . 2018 : e.T14201A22064312. doi :10.2305/IUCN.UK.2018-2.RLTS.T14201A22064312.en .
^ a b Csorba, G.; Rosell-Ambal, R. G. B.; Tabaranza, B.; Sedlock, J.; Ingle, N. R.; Heaney, L.; Balete, D. S.; Ong, P. (2016). "Myotis rufopictus " . IUCN Red List of Threatened Species . 2016 : e.T136411A22017446. doi :10.2305/IUCN.UK.2016-2.RLTS.T136411A22017446.en .
^ a b Arroyo-Cabrales, J.; Ospina-Garces, S. (2016). "Myotis peninsularis " . IUCN Red List of Threatened Species . 2016 : e.T14189A22066405. doi :10.2305/IUCN.UK.2016-1.RLTS.T14189A22066405.en .
^ a b Csorba, G.; Görföl, T. (2017). "Myotis peytoni " . IUCN Red List of Threatened Species . 2017 : e.T85568321A85568324. doi :10.2305/IUCN.UK.2017-2.RLTS.T85568321A85568324.en .
^ a b Piraccini, R. (2016). "Myotis dasycneme " . IUCN Red List of Threatened Species . 2016 : e.T14127A22055164. doi :10.2305/IUCN.UK.2016-2.RLTS.T14127A22055164.en .
^ a b Solari, S. (2019). "Myotis ruber " . IUCN Red List of Threatened Species . 2019 : e.T14197A22062092. doi :10.2305/IUCN.UK.2019-3.RLTS.T14197A22062092.en .
^ a b Ruedi, M.; Csorba, G.; Liang-Kong, L.; Cheng-Han, C. (2017). "Myotis soror " . IUCN Red List of Threatened Species . 2017 : e.T85342662A85342666. doi :10.2305/IUCN.UK.2017-2.RLTS.T85342662A85342666.en .
^ a b Csorba, G.; Görföl, T. (2020). "Myotis rufoniger " . IUCN Red List of Threatened Species . 2020 : e.T85735909A85735913. doi :10.2305/IUCN.UK.2020-3.RLTS.T85735909A85735913.en .
^ a b Azhar, M. I. (2020). "Myotis ridleyi " . IUCN Red List of Threatened Species . 2020 : e.T14194A22062376. doi :10.2305/IUCN.UK.2020-2.RLTS.T14194A22062376.en .
^ a b Barquez, R.; Perez, S.; Diaz, M. (2016). "Myotis riparius " . IUCN Red List of Threatened Species . 2016 : e.T14195A22062950. doi :10.2305/IUCN.UK.2016-1.RLTS.T14195A22062950.en .
^ a b Monadjem, A.; Jacobs, D. (2017). "Myotis bocagii " . IUCN Red List of Threatened Species . 2017 : e.T14148A22059585. doi :10.2305/IUCN.UK.2017-2.RLTS.T14148A22059585.en .
^ a b Piraccini, R. (2016). "Myotis schaubi " . IUCN Red List of Threatened Species . 2016 : e.T14198A22061746. doi :10.2305/IUCN.UK.2016-2.RLTS.T14198A22061746.en .
^ a b Larsen, R. (2016). "Myotis martiniquensis " . IUCN Red List of Threatened Species . 2016 : e.T76435251A22066280. doi :10.2305/IUCN.UK.2016-1.RLTS.T76435251A22066280.en .
^ a b Benda, P.; Lavrenchenko, L. (2017). "Myotis scotti " . IUCN Red List of Threatened Species . 2017 : e.T14199A22062198. doi :10.2305/IUCN.UK.2017-2.RLTS.T14199A22062198.en .
^ a b Zhigalin, A. (2020). "Myotis sibiricus " . IUCN Red List of Threatened Species . 2020 : e.T85567062A85567065. doi :10.2305/IUCN.UK.2020-2.RLTS.T85567062A85567065.en .
^ a b Barquez, R.; Diaz, M.; Samudio Jr, R.; Arroyo-Cabrales, J. (2016). "Myotis albescens " . IUCN Red List of Threatened Species . 2016 : e.T14140A22049892. doi :10.2305/IUCN.UK.2016-1.RLTS.T14140A22049892.en .
^ a b c Csorba, G.; Bates, P.; Lee, B.; Soisook, P. (2016). "Myotis oreias " . IUCN Red List of Threatened Species . 2016 : e.T14186A22067080. doi :10.2305/IUCN.UK.2016-2.RLTS.T14186A22067080.en .
^ a b Arroyo-Cabrales, J.; Álvarez-Castañeda, S. T. (2017). "Myotis austroriparius " . IUCN Red List of Threatened Species . 2017 : e.T14147A22059907. doi :10.2305/IUCN.UK.2017-2.RLTS.T14147A22059907.en .
^ a b Barquez, R.; Diaz, M. (2017) [errata version of 2016 assessment]. "Myotis aelleni " . IUCN Red List of Threatened Species . 2016 : e.T14139A115121458. doi :10.2305/IUCN.UK.2016-3.RLTS.T14139A22049723.en .
^ a b Arroyo-Cabrales, J.; Miller, B.; Reid, F.; Cuarón, A. D.; de Grammont, P. C. (2017). "Myotis auriculus " . IUCN Red List of Threatened Species . 2017 : e.T14145A22060698. doi :10.2305/IUCN.UK.2017-2.RLTS.T14145A22060698.en .
^ a b Jiang, T. L.; Feng, J. (2019). "Myotis altarium " . IUCN Red List of Threatened Species . 2019 : e.T14141A22050057. doi :10.2305/IUCN.UK.2019-3.RLTS.T14141A22050057.en .
^ a b Csorba, G.; Bumrungsri, S.; Bates, P. (2020). "Myotis rosseti " . IUCN Red List of Threatened Species . 2020 : e.T14196A22062800. doi :10.2305/IUCN.UK.2020-2.RLTS.T14196A22062800.en .
^ a b Barquez, R.; Diaz, M. (2016). "Myotis simus " . IUCN Red List of Threatened Species . 2016 : e.T14204A22064642. doi :10.2305/IUCN.UK.2016-2.RLTS.T14204A22064642.en .
^ a b Srinivasulu, C.; Srinivasulu, B. (2019). "Myotis muricola " . IUCN Red List of Threatened Species . 2019 : e.T85537578A22065403. doi :10.2305/IUCN.UK.2019-3.RLTS.T85537578A22065403.en .
^ a b Bouillard, N.; Csorba, G.; Görföl, T. (2021). "Myotis weberi " . IUCN Red List of Threatened Species . 2021 : e.T85736011A85736023. doi :10.2305/IUCN.UK.2021-2.RLTS.T85736011A85736023.en .
^ a b Monadjem, A.; Taylor, P. J.; Jacobs, D.; Cotterill, F. (2017). "Myotis welwitschii " . IUCN Red List of Threatened Species . 2017 : e.T14211A22068792. doi :10.2305/IUCN.UK.2017-2.RLTS.T14211A22068792.en .
^ a b Coroiu, I. (2016). "Myotis mystacinus " . IUCN Red List of Threatened Species . 2016 : e.T14134A22052250. doi :10.2305/IUCN.UK.2016-2.RLTS.T14134A22052250.en .
^ a b Fukui, D.; Sano, A. (2021) [errata version of 2019 assessment]. "Myotis yanbarensis " . IUCN Red List of Threatened Species . 2019 : e.T29484A209551473. doi :10.2305/IUCN.UK.2019-3.RLTS.T29484A209551473.en .
^ a b Barquez, R.; Diaz, M. (2017) [errata version of 2016 assessment]. "Myotis levis " . IUCN Red List of Threatened Species . 2016 : e.T14174A115121699. doi :10.2305/IUCN.UK.2016-3.RLTS.T14174A22056440.en .
^ a b Solari, S. (2019). "Myotis yumanensis " . IUCN Red List of Threatened Species . 2019 : e.T14213A22068335. doi :10.2305/IUCN.UK.2019-1.RLTS.T14213A22068335.en .
^ a b Ruedi, M.; Csorba, G.; Liang-Kong, L.; Cheng-Han, C. (2017). "Submyotodon latirostris " . IUCN Red List of Threatened Species . 2017 : e.T85537971A85537974. doi :10.2305/IUCN.UK.2017-2.RLTS.T85537971A85537974.en .
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